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Equity Incentive Awards (Tables)
 9 Months Ended
Sep. 30, 2021
Share-based Payment Arrangement [Abstract]  
Summary of Stock Options Activity The following table summarizes the Company’s option activity and related information:
Number of
Options
(in thousands)		Weighted-
Average
Exercise Price
Balance at December 31, 202010,432 $11.65 
Options granted6,863 8.02 
Options exercised(9)5.86 
Options forfeited(3,427)11.43 
Balance at September 30, 202113,859 $9.91 
Exercisable as of September 30, 20215,724 $10.11 
Summary of Restricted Stock Units Activity The following table summarizes the Company’s RSUs activity and related information:
Number of Units
(in thousands)		Weighted-
Average Grant-
Date Fair Value
Outstanding at December 31, 2020540 $6.41 
Granted2,570 3.91 
Vested and released(289)5.35 
Forfeited(228)8.95 
Outstanding at September 30, 20212,593 $3.83 
Stock-Based Compensation Expense The following table presents, by operating expense, the Company’s stock-based compensation expense:
Three Months Ended
September 30,		Nine Months Ended
September 30,
2021202020212020
(In thousands)
Research and development$2,207 $2,051 $7,125 $4,967 
General and administrative3,119 3,969 13,659 9,247 
Total stock-based compensation expense$5,326 $6,020 $20,784 $14,214