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Equity Incentive Awards
 3 Months Ended
Mar. 31, 2022
Share-based Payment Arrangement [Abstract]  
Equity Incentive Awards Equity Incentive Awards
Stock Options
The following table summarizes the Company’s option activity and related information:
Number of
Options
(in thousands)		Weighted-
Average
Exercise Price
Balance at December 31, 202113,288 $9.34 
Options granted5,735 1.40 
Options exercised(15)0.19 
Options forfeited(2,837)6.60 
Balance at March 31, 202216,171 $7.01 
Exercisable as of March 31, 20223,921 $12.95 
Options granted during the three months ended March 31, 2022 include 0.9 million shares of performance-based non-qualified options with both performance and service vesting conditions.
Restricted Stock Units (“RSUs”)
The following table summarizes the Company’s RSUs activity and related information:
Number of Units
(in thousands)		Weighted-
Average Grant-
Date Fair Value
Outstanding at December 31, 20212,126 $3.76 
Granted858 1.43 
Vested and released(371)4.62 
Forfeited(91)3.83 
Outstanding at March 31, 20222,522 $2.84 
RSUs granted during the three months ended March 31, 2022 include 0.2 million shares of performance units with both performance and service vesting conditions.
Stock-Based Compensation Expense
The following table presents, by operating expense, the Company’s stock-based compensation expense:
Three Months Ended
March 31,
20222021
(In thousands)
Research and development$1,933 $2,339 
General and administrative3,448 4,885 
Total stock-based compensation expense$5,381 $7,224