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Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($)
shares in Thousands, $ in Thousands
 3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Income Statement [Abstract]    
License revenue $ 0 $ 7,500
Operating expenses:    
Research and development 22,964 19,980
General and administrative 15,169 16,163
Total operating expenses 38,133 36,143
Operating loss (38,133) (28,643)
Other income, net 244 207
Net loss before income taxes (37,889) (28,436)
Income tax provision (19) 0
Net loss (37,908) (28,436)
Other comprehensive income (loss):    
Net unrealized loss on marketable securities (723) (22)
Foreign currency translation adjustment 14 (33)
Comprehensive loss $ (38,617) $ (28,491)
Net loss per share — basic (in USD per share) $ (0.38) $ (0.29)
Net loss per share — diluted (in USD per share) $ (0.38) $ (0.29)
Weighted-average common shares used to compute net loss per share - basic (in shares) 98,596 97,750
Weighted-average common shares used to compute net loss per share - diluted (in shares) 98,596 97,750