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Condensed Consolidated Statements of Cash Flows - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Cash flows from operating activities:    
Net loss $ (37,908) $ (28,436)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 1,425 1,166
Stock-based compensation expense 5,381 7,224
Amortization of premium and accrued interest on marketable securities 212 855
Other (92) (42)
Changes in operating assets and liabilities:    
Prepaid expenses and other current assets (2,916) (1,279)
Other assets (1,658) (464)
Operating lease right-of-use asset 1,126 332
Accounts payable 1,437 (763)
Accrued expenses and other current liabilities (1,207) (396)
Other non-current liabilities 19 0
Lease liability and lease incentive receivable 5,252 (502)
Net cash used in operating activities (28,929) (22,305)
Cash flows from investing activities:    
Purchases of marketable securities (29,750) (82,667)
Maturities of marketable securities 118,507 90,650
Purchases of property and equipment (4,374) (1,404)
Net cash provided by investing activities 84,383 6,579
Cash flows from financing activities:    
Proceeds from offerings of common stock, net of issuance costs 0 1,694
Proceeds from issuance of common stock pursuant to option exercises 3 51
Repayment of loan 0 (240)
Net cash provided by financing activities 3 1,505
Net increase (decrease) in cash and cash equivalents and restricted cash 55,457 (14,221)
Cash and cash equivalents and restricted cash at beginning of period 36,698 63,423
Cash and cash equivalents and restricted cash at end of period 92,155 49,202
Cash and cash equivalents 89,652 45,422
Restricted cash 2,503 3,780
Cash and cash equivalents and restricted cash at end of period 92,155 49,202
Supplemental schedule of noncash investing and financing information    
Fixed assets in accounts payable, accrued expenses and other current liabilities $ 2,616 $ 805