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Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($)
shares in Thousands, $ in Thousands
 3 Months Ended 6 Months Ended
Jun. 30, 2022
Jun. 30, 2021
Jun. 30, 2022
Jun. 30, 2021
Income Statement [Abstract]        
License revenue $ 0 $ 0 $ 0 $ 7,500
Operating expenses:        
Research and development 30,265 22,608 53,229 42,588
General and administrative 13,760 21,930 28,929 38,093
Total operating expenses 44,025 44,538 82,158 80,681
Operating loss (44,025) (44,538) (82,158) (73,181)
Other income, net 283 205 527 412
Net loss before income taxes (43,742) (44,333) (81,631) (72,769)
Income tax provision (19) 0 (38) 0
Net loss (43,761) (44,333) (81,669) (72,769)
Other comprehensive loss:        
Net unrealized loss on marketable securities (339) (62) (1,062) (84)
Foreign currency translation adjustment (34) (39) (20) (72)
Comprehensive loss $ (44,134) $ (44,434) $ (82,751) $ (72,925)
Net loss per share — basic (in USD per share) $ (0.44) $ (0.45) $ (0.83) $ (0.74)
Net loss per share — diluted (in USD per share) $ (0.44) $ (0.45) $ (0.83) $ (0.74)
Weighted-average common shares used to compute net loss per share - basic (in shares) 99,000 98,018 98,799 97,885
Weighted-average common shares used to compute net loss per share - diluted (in shares) 99,000 98,018 98,799 97,885