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Condensed Consolidated Statements of Cash Flows - USD ($)
$ in Thousands
 6 Months Ended
Jun. 30, 2022
Jun. 30, 2021
Cash flows from operating activities:    
Net loss $ (81,669) $ (72,769)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 3,028 2,361
Stock-based compensation expense 10,323 15,458
Amortization of premium and accrued interest on marketable securities 258 1,398
Loss on disposal of property and equipment 101 0
Impairment of long-lived assets 2,011 0
Other (76) (81)
Changes in operating assets and liabilities:    
Prepaid expenses and other current assets (4,165) (1,653)
Other assets (3,403) (99)
Operating lease right-of-use asset 2,234 1,505
Accounts payable 3,216 (394)
Accrued expenses and other current liabilities 2,172 (583)
Other non-current liabilities (38) 0
Lease liability and lease incentive receivable 8,200 (1,363)
Net cash used in operating activities (57,808) (56,220)
Cash flows from investing activities:    
Purchases of marketable securities (57,674) (199,959)
Maturities of marketable securities 183,599 240,377
Sales of marketable securities 0 8,990
Purchases of property and equipment (10,807) (5,052)
Net cash provided by investing activities 115,118 44,356
Cash flows from financing activities:    
Proceeds from offerings of common stock, net of issuance costs 0 1,685
Proceeds from issuance of common stock pursuant to option exercises 3 51
Proceeds from employee stock purchase plan 362 526
Repayment of loan 0 (240)
Net cash provided by financing activities 365 2,022
Net increase (decrease) in cash and cash equivalents and restricted cash 57,675 (9,842)
Cash and cash equivalents and restricted cash at beginning of period 36,698 63,423
Cash and cash equivalents and restricted cash at end of period 94,373 53,581
Cash and cash equivalents 91,870 47,299
Restricted cash 2,503 6,282
Cash and cash equivalents and restricted cash at end of period 94,373 53,581
Supplemental schedule of noncash investing and financing information    
Right-of-use assets obtained in exchange for lease liability 0 84,005
Fixed assets in accounts payable, accrued expenses and other current liabilities $ 908 $ 1,562