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Condensed Consolidated Statements of Operations and Comprehensive Loss - USD ($)
shares in Thousands, $ in Thousands
 3 Months Ended 9 Months Ended
Sep. 30, 2022
Sep. 30, 2021
Sep. 30, 2022
Sep. 30, 2021
Income Statement [Abstract]        
License revenue $ 0 $ 0 $ 0 $ 7,500
Operating expenses:        
Research and development 23,849 24,069 77,078 66,657
General and administrative 17,188 14,453 46,117 52,546
Total operating expenses 41,037 38,522 123,195 119,203
Operating loss (41,037) (38,522) (123,195) (111,703)
Other income, net 923 160 1,450 572
Net loss before income taxes (40,114) (38,362) (121,745) (111,131)
Income tax provision (17) 0 (55) 0
Net loss (40,131) (38,362) (121,800) (111,131)
Other comprehensive loss:        
Net unrealized loss on marketable securities (127) (55) (1,189) (139)
Foreign currency translation adjustment (30) (51) (50) (123)
Comprehensive loss $ (40,288) $ (38,468) $ (123,039) $ (111,393)
Net loss per share — basic (in USD per share) $ (0.40) $ (0.39) $ (1.23) $ (1.13)
Net loss per share — diluted (in USD per share) $ (0.40) $ (0.39) $ (1.23) $ (1.13)
Weighted-average common shares used to compute net loss per share - basic (in shares) 99,475 98,126 99,027 97,966
Weighted-average common shares used to compute net loss per share - diluted (in shares) 99,475 98,126 99,027 97,966