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Equity Incentive Awards
 6 Months Ended
Jun. 30, 2023
Share-Based Payment Arrangement [Abstract]  
Equity Incentive Awards Equity Incentive Awards
Stock Options
The following table summarizes the Company’s option activity and related information:
Number of
Options
(in thousands)		Weighted-
Average
Exercise Price
Balance at December 31, 202219,320 $5.79 
Options granted4,736 0.81 
Options exercised— — 
Options forfeited(541)5.13 
Balance at June 30, 202323,515 $4.80 
Exercisable as of June 30, 20238,867 $8.82 
Restricted Stock Units (“RSUs”)
The following table summarizes the Company’s RSUs activity and related information:
Number of Units
(in thousands)		Weighted-
Average Grant-
Date Fair Value
Outstanding at December 31, 20221,693 $2.90 
Granted534 0.76 
Vested and released(458)3.25 
Forfeited(44)3.14 
Outstanding at June 30, 20231,725 $2.14 
Stock-Based Compensation Expense
The following table presents, by operating expense, the Company’s stock-based compensation expense:
Three Months Ended
June 30,		Six Months Ended
June 30,
2023202220232022
(In thousands)
Research and development$1,380 $1,830 $2,761 $3,763 
General and administrative3,089 3,112 6,271 6,560 
Total stock-based compensation expense$4,469 $4,942 $9,032 $10,323