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Equity Incentive Awards
 9 Months Ended
Sep. 30, 2023
Share-Based Payment Arrangement [Abstract]  
Equity Incentive Awards Equity Incentive Awards
Stock Options
The following table summarizes the Company’s option activity and related information:
Number of
Options
(in thousands)		Weighted-
Average
Exercise Price
Balance at December 31, 202219,320 $5.79 
Options granted6,056 1.05 
Options exercised(1)1.29 
Options forfeited(1,314)3.05 
Balance at September 30, 202324,061 $4.74 
Exercisable as of September 30, 20239,724 $8.54 
Restricted Stock Units (“RSUs”)
The following table summarizes the Company’s RSUs activity and related information:
Number of Units
(in thousands)		Weighted-
Average Grant-
Date Fair Value
Outstanding at December 31, 20221,693 $2.90 
Granted539 0.77 
Vested and released(491)3.14 
Forfeited(128)1.94 
Outstanding at September 30, 20231,613 $2.20 
Stock-Based Compensation Expense
The following table presents, by operating expense, the Company’s stock-based compensation expense:
Three Months Ended
September 30,		Nine Months Ended
September 30,
2023202220232022
(In thousands)
Research and development$1,164 $1,432 $3,925 $5,195 
General and administrative3,225 3,071 9,496 9,631 
Total stock-based compensation expense$4,389 $4,503 $13,421 $14,826