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Organization and Summary of Significant Accounting Policies - Additional Information (Detail) - USD ($)	12 Months Ended
	Dec. 31, 2021	Dec. 31, 2020	Dec. 31, 2019
Property, Plant and Equipment [Line Items]
Year founded	2010
Impairment of long-lived assets	$ 0
Accumulated deficit	(196,092,000)	$ (166,893,000)
Net loss	(29,199,000)	(44,836,000)	$ (39,612,000)
Equipment Financing Leases	499,000	393,000	$ 388,000
Current Liabilities	4,854,000	$ 6,945,000
ATM Facility [Member]
Property, Plant and Equipment [Line Items]
Aggregate common stock offering value	$ 50,000,000.0
Cellectis [Member]
Property, Plant and Equipment [Line Items]
Percentage of ownership in outstanding common stock	61.80%

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