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Income Taxes - Deferred Tax Assets and Liabilities (Details) - USD ($) $ in Thousands | Dec. 31, 2021 | Dec. 31, 2020
--- | --- | ---
Deferred tax assets: |  |
Net operating loss and credit carryforwards | $ 107,979 | $ 104,301
Operating lease liability | 1,878 | 2,394
Accrued bonus | 981 | 850
Accrued expenses | 1,566 | 429
Deferred revenue | 309 | 1,018
Equity method investment | 1,243 | 1,031
Other | 925 | 394
Gross deferred tax asset | 114,881 | 110,417
Valuation allowance | (113,276) | (108,300)
Net deferred tax asset | 1,605 | 2,117
Deferred tax liabilities: |  |
Operating lease asset | (1,429) | (1,941)
Other | (176) | (176)
Total deferred tax liabilities | (1,605) | (2,117)
Net deferred income tax assets and liabilities: | $ 0 | $ 0

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