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Collaboration and License Agreements - Additional Information (Details) - USD ($)			6 Months Ended
	Sep. 10, 2018	Feb. 24, 2016	Jun. 30, 2019	Jun. 30, 2018	Jan. 01, 2019	Dec. 31, 2018
Collaboration and License Agreements [Line items]
Deferred revenue			$ 7,129,000		$ 8,029,000	$ 8,436,000
Development and Commercial License Agreement | Servier
Collaboration and License Agreements [Line items]
Upfront payment		$ 105,000,000
Recognized revenue under agreement			3,000,000	$ 2,900,000
Deferred revenue			85,200,000			88,600,000
Development or sales-based milestone payments			0	$ 0
Development and Commercial License Agreement | Servier | Current Liabilities
Collaboration and License Agreements [Line items]
Deferred revenue			5,900,000			6,400,000
Development and Commercial License Agreement | Servier | Maximum
Collaboration and License Agreements [Line items]
Total payments to be received		1,600,000,000
Aggregate milestone payments		1,500,000,000
Development milestone payments		401,300,000
Commercial milestone payments		$ 1,100,000,000
Collaboration and License Agreement | Gilead
Collaboration and License Agreements [Line items]
Recognized revenue under agreement			6,700,000
Deferred revenue			1,200,000			2,300,000
Development or sales-based milestone payments			$ 0
Research funding payment term	3 years		3 years
Semi-annual research funding payments	$ 40,000,000
Collaboration and License Agreement | Gilead | Current Liabilities
Collaboration and License Agreements [Line items]
Deferred revenue			$ 1,200,000			$ 2,300,000
Collaboration and License Agreement | Gilead | Maximum
Collaboration and License Agreements [Line items]
Development milestone payments	105,000,000
Commercial milestone payments	340,000,000
Receivable from research funding	40,000,000
Development and commercial milestone payments	$ 445,000,000

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- Definition

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- Definition

Commercial milestone payments.

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- Definition

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- Definition

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- Definition

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