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Segment Reporting - Financial Information with Respect to Company's Reportable Segments (Details) - USD ($) $ in Thousands | 3 Months Ended | 9 Months Ended
--- | --- | ---
Sep. 30, 2019 | Sep. 30, 2018 | Sep. 30, 2019 | Sep. 30, 2018
Segment Reporting Information [Line Items] |  |  |  |
Revenue | $ 4,865 | $ 2,541 | $ 15,716 | $ 5,943
Total allocation of centralized research and development operational cash expenditures | 19,791 | 9,737 | 62,512 | 28,723
Loss from operations | (21,978) | (10,447) | (65,343) | (31,807)
Adjustments to reconcile segment operating loss to consolidated loss from operations: |  |  |  |
Interest income received | (1,236) | (691) | (3,322) | (1,213)
Depreciation and amortization |  |  | (3,508) | (1,555)
Share-based compensation |  |  | (6,319) | (697)
Loss on disposal of assets |  |  | (22) | (8)
Changes in prepaid expenses, accounts payable and accrued expenses | 5,179 | 7,866 | 15,988 | 10,772
Operating Segments |  |  |  |
Segment Reporting Information [Line Items] |  |  |  |
Revenue | 4,865 | 2,541 | 15,716 | 5,943
Total segment operational cash expenditures | 16,232 | 12,584 | 48,198 | 28,493
Total allocation of centralized research and development operational cash expenditures | 4,081 | 2,984 | 10,952 | 8,159
Loss from operations | (15,448) | (13,027) | (43,434) | (30,709)
Operating Segments | Therapeutics |  |  |  |
Segment Reporting Information [Line Items] |  |  |  |
Revenue | 4,719 | 2,115 | 14,357 | 5,009
Total segment operational cash expenditures | 14,290 | 11,727 | 42,072 | 25,903
Total allocation of centralized research and development operational cash expenditures | 4,081 | 2,387 | 10,952 | 6,329
Loss from operations | (13,652) | (11,999) | (38,667) | (27,223)
Operating Segments | Food |  |  |  |
Segment Reporting Information [Line Items] |  |  |  |
Revenue | 146 | 426 | 1,359 | 934
Total segment operational cash expenditures | 1,942 | 857 | 6,126 | 2,590
Total allocation of centralized research and development operational cash expenditures |  | 597 |  | 1,830
Loss from operations | (1,796) | (1,028) | (4,767) | (3,486)
Segment Reconciling Items |  |  |  |
Adjustments to reconcile segment operating loss to consolidated loss from operations: |  |  |  |
Corporate general and administrative cash expenditures | (6,581) | (3,719) | (24,726) | (8,397)
Interest income received | (1,236) | (691) | (3,322) | (1,213)
Depreciation and amortization | (1,401) | (584) | (3,508) | (1,555)
Share-based compensation | $ (2,491) | $ (292) | (6,319) | (697)
Loss on disposal of assets |  |  | $ (22) | $ (8)

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