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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS (Unaudited) - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Cash flows from operating activities:    
Net loss $ (28,168) $ (18,689)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 1,968 2,176
Share-based compensation 4,390 3,632
Loss on disposal of assets 11 23
Non-cash interest expense 17  
Amortization of right-of-use assets 288 290
Loss from equity method investments 952  
Amortization of discount on note receivable (87)  
Changes in operating assets and liabilities:    
Prepaid expenses 374 (619)
Accounts receivable   (9,263)
Other assets and other current assets 171 (6)
Accounts payable (374) 221
Other liabilities and other current liabilities (5,150) (2,146)
Deferred revenue (2,830) 92,914
Lease liabilities and right-of-use assets (434) (461)
Net cash provided by (used in) operating activities (28,872) 68,072
Cash flows from investing activities:    
Purchases of property, equipment and software (818) (1,125)
Net cash used in investing activities (818) (1,125)
Cash flows from financing activities:    
Proceeds from stock option exercises 340 1,297
Proceeds from employee stock purchase plan 282 418
Proceeds from issuance of common stock to collaboration partners   35,000
Proceeds from offering of common stock, net of issuance costs 1,627  
Net cash provided by financing activities 2,249 36,715
Net increase (decrease) in cash and cash equivalents (27,441) 103,662
Cash and cash equivalents—beginning of period 143,663 89,798
Cash and cash equivalents —end of period 116,222 193,460
Supplemental disclosures of noncash activities:    
Property, equipment and software additions included in accounts payable and other current liabilities 74 $ 637
Unsettled at-the-market issuances of common stock included in other current assets $ 217