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CONDENSED CONSOLIDATED STATEMENTS OF CHANGES IN STOCKHOLDERS' EQUITY (Unaudited) - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-in Capital | Accumulated Deficit | Treasury Stock
--- | --- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2020 | $ 44,425 |  | $ 331,450 | $ (286,073) | $ (952)
Beginning balance, Shares at Dec. 31, 2020 |  | 53,503,124 |  |  |
Stock option exercises | 1,297 |  | 1,297 |  |
Stock option exercises, Shares |  | 676,791 |  |  |
Issuance of common stock under employee stock purchase plan | 418 |  | 418 |  |
Issuance of common stock under employee stock purchase plan, Shares |  | 90,796 |  |  |
Share-based compensation expense | 3,632 |  | 3,632 |  |
Issuance of common stock to collaboration partners | 27,739 |  | 27,739 |  |
Issuance of common stock to collaboration partners, Shares |  | 3,762,190 |  |  |
Net income (loss) | (18,689) |  |  | (18,689) |
Ending balance at Mar. 31, 2021 | 58,822 |  | 364,536 | (304,762) | (952)
Ending balance, Shares at Mar. 31, 2021 |  | 58,032,901 |  |  |
Beginning balance at Dec. 31, 2020 | 44,425 |  | 331,450 | (286,073) | (952)
Beginning balance, Shares at Dec. 31, 2020 |  | 53,503,124 |  |  |
Net income (loss) | (8,312) |  |  |  |
Ending balance at Sep. 30, 2021 | 101,443 |  | 396,780 | (294,385) | (952)
Ending balance, Shares at Sep. 30, 2021 |  | 60,892,018 |  |  |
Beginning balance at Mar. 31, 2021 | 58,822 |  | 364,536 | (304,762) | (952)
Beginning balance, Shares at Mar. 31, 2021 |  | 58,032,901 |  |  |
Stock option exercises | 1,353 |  | 1,353 |  |
Stock option exercises, Shares |  | 766,767 |  |  |
Share-based compensation expense | 3,896 |  | 3,896 |  |
Proceeds from issuance of common stock, net of issuance cost | 14,826 |  | 14,826 |  |
Proceeds from issuance of common stock, net of issuance cost, Shares |  | 1,293,754 |  |  |
Net income (loss) | 21,656 |  |  | 21,656 |
Ending balance at Jun. 30, 2021 | 100,553 |  | 384,611 | (283,106) | (952)
Ending balance, Shares at Jun. 30, 2021 |  | 60,093,422 |  |  |
Stock option exercises | 3,393 |  | 3,393 |  |
Stock option exercises, Shares |  | 414,073 |  |  |
Issuance of common stock under employee stock purchase plan | 386 |  | 386 |  |
Issuance of common stock under employee stock purchase plan, Shares |  | 36,091 |  |  |
Share-based compensation expense | 4,551 |  | 4,551 |  |
Proceeds from issuance of common stock, net of issuance cost | 3,839 |  | 3,839 |  |
Proceeds from issuance of common stock, net of issuance cost, Shares |  | 348,432 |  |  |
Net income (loss) | (11,279) |  |  | (11,279) |
Ending balance at Sep. 30, 2021 | 101,443 |  | 396,780 | (294,385) | (952)
Ending balance, Shares at Sep. 30, 2021 |  | 60,892,018 |  |  |
Beginning balance at Dec. 31, 2021 | $ 91,168 |  | 408,795 | (316,675) | (952)
Beginning balance, Shares at Dec. 31, 2021 | 61,712,577 | 61,712,577 |  |  |
Stock option exercises | $ 340 |  | 340 |  |
Stock option exercises, Shares |  | 233,001 |  |  |
Issuance of common stock under employee stock purchase plan | 282 |  | 282 |  |
Issuance of common stock under employee stock purchase plan, Shares |  | 78,060 |  |  |
Share-based compensation expense | 4,390 |  | 4,390 |  |
Proceeds from issuance of common stock, net of issuance cost | 1,647 |  | 1,647 |  |
Proceeds from issuance of common stock, net of issuance cost, Shares |  | 561,405 |  |  |
Net income (loss) | (28,168) |  |  | (28,168) |
Ending balance at Mar. 31, 2022 | 69,659 |  | 415,454 | (344,843) | (952)
Ending balance, Shares at Mar. 31, 2022 |  | 62,585,043 |  |  |
Beginning balance at Dec. 31, 2021 | $ 91,168 |  | 408,795 | (316,675) | (952)
Beginning balance, Shares at Dec. 31, 2021 | 61,712,577 | 61,712,577 |  |  |
Stock option exercises, Shares | 330,359 |  |  |  |
Net income (loss) | $ (83,149) |  |  |  |
Ending balance at Sep. 30, 2022 | $ 84,380 | $ 1 | 485,155 | (399,824) | (952)
Ending balance, Shares at Sep. 30, 2022 | 111,745,219 | 111,745,219 |  |  |
Beginning balance at Mar. 31, 2022 | $ 69,659 |  | 415,454 | (344,843) | (952)
Beginning balance, Shares at Mar. 31, 2022 |  | 62,585,043 |  |  |
Stock option exercises | 38 |  | 38 |  |
Stock option exercises, Shares |  | 78,234 |  |  |
Share-based compensation expense | 5,138 |  | 5,138 |  |
Proceeds from issuance of common stock, net of issuance cost | 47,670 |  | 47,670 |  |
Proceeds from issuance of common stock, net of issuance cost, Shares |  | 36,274,651 |  |  |
Issuance of common stock to collaboration partners | 11,553 |  | 11,553 |  |
Issuance of common stock to collaboration partners, Shares |  | 12,407,440 |  |  |
Restricted stock units vested, Shares |  | 260,137 |  |  |
Net income (loss) | (31,035) |  |  | (31,035) |
Ending balance at Jun. 30, 2022 | 103,023 |  | 479,853 | (375,878) | (952)
Ending balance, Shares at Jun. 30, 2022 |  | 111,605,505 |  |  |
Stock option exercises | 9 |  | 9 |  |
Stock option exercises, Shares |  | 19,124 |  |  |
Issuance of common stock under employee stock purchase plan | 161 | $ 1 | 160 |  |
Issuance of common stock under employee stock purchase plan, Shares |  | 116,612 |  |  |
Share-based compensation expense | 5,133 |  | 5,133 |  |
Restricted stock units vested, Shares |  | 3,978 |  |  |
Net income (loss) | (23,946) |  |  | (23,946) |
Ending balance at Sep. 30, 2022 | $ 84,380 | $ 1 | $ 485,155 | $ (399,824) | $ (952)
Ending balance, Shares at Sep. 30, 2022 | 111,745,219 | 111,745,219 |  |  |

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