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Collaboration and License Agreements - Additional Information (Details) - USD ($) |  |  |  | 1 Months Ended | 3 Months Ended | 9 Months Ended |  |
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Jun. 14, 2022 | Apr. 09, 2021 | Jan. 06, 2021 | Jul. 31, 2022 | Sep. 30, 2022 | Sep. 30, 2021 | Sep. 30, 2022 | Sep. 30, 2021 | Dec. 31, 2021 | Aug. 31, 2021
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Equity method investments |  |  |  |  |  |  |  |  | $ 3,751,000 |
Revenue |  |  |  |  | $ 7,363,000 | $ 24,036,000 | $ 14,500,000 | $ 109,190,000 |  |
Deferred revenue |  |  |  |  | $ 48,405,000 |  | $ 48,405,000 |  | $ 21,244,000 |
Common stock, shares issued |  |  |  |  | 111,745,219 |  | 111,745,219 |  | 61,712,577 |
Proceeds from offering of common stock, net of issuance costs |  |  |  |  |  |  | $ 49,345,000 | 18,130,000 |  |
Prepaid expenses |  |  |  |  | $ 12,292,000 |  | 12,292,000 |  | $ 17,434,000 |
Research and development |  |  |  |  | 19,959,000 | 25,940,000 | 62,867,000 | 88,768,000 |  |
Revenue recorded included in deferred revenue |  |  |  |  |  |  | 10,800,000 |  |  |
Servier | Program Purchase Agreement |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Research and development |  | $ 1,250,000 |  |  |  |  |  |  |  |
Waiver of milestone payment |  | $ 18,750,000 |  |  |  |  |  |  |  |
Research and Development Arrangement |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Prepaid Expense and Other Assets |  |  |  |  | 11,200,000 |  | 11,200,000 |  |  |
Prepaid expenses |  |  |  |  | 4,200,000 |  | 4,200,000 |  |  |
Other Assets |  |  |  |  | 7,000,000.0 |  | 7,000,000.0 |  |  |
Research and development |  |  |  |  |  |  | 1,700,000 |  |  |
Research and Development Arrangement | Current Liabilities |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  | 30,100,000 |  | 30,100,000 |  |  |
Research and Development Arrangement | Novartis Pharma AG |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Upfront cash payment | $ 50,000,000.0 |  |  |  |  |  |  |  |  |
Milestone payments to be received for additional targets | $ 1,400,000,000 |  |  |  |  |  |  |  |  |
Obligation to pay royalties expiry date | 10 years |  |  |  |  |  |  |  |  |
Transaction price from the Stock Purchase Agreement | $ 13,400,000 |  |  |  |  |  |  |  |  |
Revenue |  |  |  |  | 3,600,000 |  | 3,600,000 |  |  |
Deferred revenue related to agreement |  |  |  |  | 60,000,000.0 |  | $ 60,000,000.0 |  |  |
Research and Development Arrangement | Novartis Pharma AG | Stock Purchase Agreement |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Upfront cash payment |  |  |  | $ 50,000,000.0 |  |  |  |  |  |
Sale of stock, number of shares issued in transaction | 12,407,440 |  |  |  |  |  |  |  |  |
Aggregate purchase price | $ 25,000,000.0 |  |  |  |  |  |  |  |  |
Sale of stock, price per share | $ 2.01 |  |  |  |  |  |  |  |  |
Percentage of premium over the volume-weighted-average-price of the company’s common stock | 20.00% |  |  |  |  |  |  |  |  |
Upfront Compensation | $ 75,000,000.0 |  |  |  |  |  |  |  |  |
Equity upon the issuance of the shares | 11,600,000 |  |  |  |  |  |  |  |  |
Transaction price allocated to the revenue arrangement | $ 63,400,000 |  |  |  |  |  |  |  |  |
Sale of stock, description of transaction |  |  |  |  |  |  | Pursuant to the Novartis Stock Purchase Agreement, subject to certain exceptions, Novartis may not sell the Novartis Shares without the Company’s approval for a period of two years following the Novartis Effective Date. In addition, for a period of two years following the Novartis Effective Date, Novartis and its affiliates may not (a) effect or otherwise participate in, directly or indirectly, any acquisition of any of our securities or material assets, any tender offer or exchange offer, merger or other business combination or change of control involving the Company, any recapitalization, restructuring, liquidation, dissolution or other extraordinary transaction with respect to the Company, or any solicitation of proxies or consents to vote any of the Company’s securities or (b) act with any other person, or publicly disclose any intention, to do any of the foregoing. The Novartis Stock Purchase Agreement also contains customary representations, warranties, and covenants of both parties. |  |  |
Research and Development Arrangement | Eli Lilly and Company |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Upfront cash payment |  |  | $ 100,000,000.0 |  |  |  |  |  |  |
Milestone payments to be received for additional targets |  |  | 420,000,000.0 |  |  |  |  |  |  |
Transaction price from the Stock Purchase Agreement |  |  | 7,300,000 |  |  |  |  |  |  |
Revenue |  |  |  |  | 3,800,000 | $ 4,900,000 | $ 10,900,000 | 15,600,000 |  |
Deferred revenue related to agreement |  |  |  |  | 78,900,000 |  | 78,900,000 |  | 88,300,000 |
Research and Development Arrangement | Eli Lilly and Company | Current Liabilities |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  | 18,300,000 |  | 18,300,000 |  | 21,200,000 |
Research and Development Arrangement | Eli Lilly and Company | Stock Purchase Agreement |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Upfront Compensation |  |  | 135,000,000.0 |  |  |  |  |  |  |
Equity upon the issuance of the shares |  |  | 27,700,000 |  |  |  |  |  |  |
Transaction price allocated to the revenue arrangement |  |  | $ 107,300,000 |  |  |  |  |  |  |
Common stock, shares issued |  |  | 3,762,190 |  |  |  |  |  |  |
Proceeds from offering of common stock, net of issuance costs |  |  | $ 35,000,000.0 |  |  |  |  |  |  |
Research and Development Arrangement | iECURE |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Investment in equity securities |  |  |  |  | 0 |  | 0 |  |  | $ 500,000
Prepaid Expense and Other Assets |  |  |  |  |  |  |  |  |  | $ 17,400,000
Development And Commercial License Agreement | Servier |  |  |  |  |  |  |  |  |  |
Collaboration and License Agreements [Line items] |  |  |  |  |  |  |  |  |  |
Deferred revenue related to agreement |  |  |  |  | $ 0 |  | 0 |  | $ 0 |
Revenue recorded included in deferred revenue |  |  |  |  |  |  | $ 0 | $ 72,900,000 |  |

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