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Condensed Consolidated Statements of Operations (Parenthetical) (Unaudited) - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2016
Mar. 31, 2015
License and milestone revenue from a related party $ 19,738 $ 11,506
Collaboration services and other revenue from a related party 8,544 4,792
Astellas Agreement [Member]    
License and milestone revenue from a related party 3,313 4,692
Collaboration services and other revenue from a related party $ 352 $ 634