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Condensed Consolidated Statements of Operations (Parenthetical) (Unaudited) - USD ($)
$ in Thousands
3 Months Ended 6 Months Ended
Jun. 30, 2016
Jun. 30, 2015
Jun. 30, 2016
Jun. 30, 2015
License and milestone revenue from a related party $ 73,197 $ 106,879 $ 92,935 $ 118,385
Collaboration services and other revenue from a related party 16,083 13,671 24,628 18,463
Astellas Agreement [Member]        
License and milestone revenue from a related party 13,043 4,860 16,356 9,552
Collaboration services and other revenue from a related party $ 326 $ 719 $ 678 $ 1,353