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Condensed Consolidated Statements of Operations (Unaudited) - USD ($) shares in Thousands, $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2016	Sep. 30, 2015	Sep. 30, 2016	Sep. 30, 2015
Revenue:
License and milestone revenue (includes $4,370, $5,120, $20,727 and $14,672 from a related party)	$ 20,867	$ 13,045	$ 113,802	$ 131,430
Collaboration services and other revenue (includes $436, $868, $1,114 and $2,221 from a related party)	9,235	6,493	33,863	24,956
Total revenue	30,102	19,538	147,665	156,386
Operating expenses:
Research and development	40,558	52,071	136,599	154,165
General and administrative	11,646	11,237	33,440	31,399
Total operating expenses	52,204	63,308	170,039	185,564
Loss from operations	(22,102)	(43,770)	(22,374)	(29,178)
Interest and other, net
Interest expense	(2,760)	(2,758)	(7,975)	(8,278)
Interest income and other, net	866	1,458	2,411	3,008
Total interest and other, net	(1,894)	(1,300)	(5,564)	(5,270)
Loss before income taxes	(23,996)	(45,070)	(27,938)	(34,448)
Provision for (benefit from) income taxes	158	28	(260)	(38)
Net loss	$ (24,154)	$ (45,098)	$ (27,678)	$ (34,410)
Net loss per share - basic and diluted	$ (0.38)	$ (0.74)	$ (0.44)	$ (0.57)
Weighted average number of common shares used to calculate net loss per share - basic and diluted	62,858	60,767	62,543	59,926

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