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Condensed Consolidated Statements of Operations (Unaudited) - USD ($) shares in Thousands, $ in Thousands	3 Months Ended
	Mar. 31, 2017	Mar. 31, 2016
Revenue:
License and milestone revenue (includes $3,249 and $3,313 from a related party)	$ 19,581	$ 19,738
Collaboration services and other revenue (includes $340 and $352 from a related party)	7,310	8,544
Total revenue	26,891	28,282
Operating expenses:
Research and development	46,732	43,650
General and administrative	11,530	11,417
Total operating expenses	58,262	55,067
Loss from operations	(31,371)	(26,785)
Interest and other, net
Interest expense	(2,375)	(2,777)
Interest income and other, net	645	1,416
Total interest and other, net	(1,730)	(1,361)
Loss before income taxes	(33,101)	(28,146)
Provision for (benefit from) income taxes	60	(305)
Net loss	$ (33,161)	$ (27,841)
Net loss per share - basic and diluted	$ (0.52)	$ (0.45)
Weighted average number of common shares used to calculate net loss per share - basic and diluted	64,037	62,184

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