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Condensed Consolidated Statements of Operations (Parenthetical) (Unaudited) - USD ($)
$ in Thousands
3 Months Ended
Mar. 31, 2017
Mar. 31, 2016
License and milestone revenue from a related party $ 19,581 $ 19,738
Collaboration services and other revenue from a related party 7,310 8,544
Astellas Agreement [Member]    
License and milestone revenue from a related party 3,249 3,313
Collaboration services and other revenue from a related party $ 340 $ 352