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Condensed Consolidated Statements of Operations (Parenthetical) (Unaudited) - USD ($)
$ in Thousands
3 Months Ended 6 Months Ended
Jun. 30, 2017
Jun. 30, 2016
Jun. 30, 2017
Jun. 30, 2016
License and milestone revenue from a related party $ 21,352 $ 73,197 $ 40,933 $ 92,935
Collaboration services and other revenue from a related party 7,645 16,083 14,955 24,628
Astellas Agreement [Member]        
License and milestone revenue from a related party 4,278 13,043 7,527 16,356
Collaboration services and other revenue from a related party $ 445 $ 326 $ 785 $ 678