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Condensed Consolidated Statements of Operations (Unaudited) (Parenthetical) - Astellas Agreement [Member] - USD ($)
$ in Thousands
3 Months Ended 9 Months Ended
Sep. 30, 2018
Sep. 30, 2017
Sep. 30, 2018
Sep. 30, 2017
License and milestone revenue from a related party $ 0 $ 0 $ 14,323 $ 0
Collaboration services and other revenue from a related party $ 5,131 $ 5,322 $ 16,448 $ 15,106