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Condensed Consolidated Statements of Operations (Unaudited) (Parenthetical) - Astellas Agreement [Member] - USD ($)
$ in Thousands
3 Months Ended 6 Months Ended
Jun. 30, 2019
Jun. 30, 2018
Jun. 30, 2019
Jun. 30, 2018
Collaboration services and other revenue from a related party $ 17,223 $ 6,145 $ 22,082 $ 11,317
License and milestone revenue from a related party $ 117,470 $ 14,323 $ 117,470 $ 14,323