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Collaboration Agreement - Related Party (Details) - USD ($) $ in Thousands | 12 Months Ended
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Dec. 31, 2021 | Dec. 31, 2020
Related Party Transaction [Line Items] |  |
Collaboration revenue - related party | $ 30,017 | $ 9,934
Takeda | Investor | Collaborative Arrangement |  |
Related Party Transaction [Line Items] |  |
Proceeds from collaborators | 3,600 | $ 14,000
Collaboration revenue - related party | $ 82,000 |

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