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Discontinued Operations - Additional Information (Details) - USD ($) $ in Thousands |  |  |  | 1 Months Ended | 3 Months Ended | 12 Months Ended |
--- | --- | --- | --- | --- | --- | --- | ---
Jul. 21, 2015 | Jul. 17, 2015 | May 04, 2015 | May 31, 2014 | Jun. 30, 2015 | Dec. 31, 2016 | Dec. 31, 2015 | Dec. 31, 2014 | Jul. 16, 2015
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
Impairment charges recorded |  |  |  |  |  | $ 0 | $ 586 | $ 0 |
Consideration in escrow |  |  |  |  |  | 0 | 500 |  |
Research and development |  |  |  |  |  | 20,076 | $ 16,440 | $ 8,287 |
Continuing Operations |  |  |  |  |  |  |  |  |
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
Research and development |  |  |  |  |  | $ 3,131 |  |  |
Research and Development Services |  |  |  |  |  |  |  |  |
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
Contract initial term |  | 18 months |  |  |  |  |  |  |
Recorded unconditional purchase obligation |  | $ 3,300 |  |  |  |  |  |  |
Affiliate |  |  |  |  |  |  |  |  |
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
General and administrative expense |  |  |  | $ 500 |  |  |  |  |
Investor | Merial | Sanofi |  |  |  |  |  |  |  |  |
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
Ownership percentage of a related party in a customer of the company |  |  |  |  |  | 100.00% |  |  |
Investor | Customer Concentration Risk | Merial | Sales |  |  |  |  |  |  |  |  |
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
Percentage of total revenue |  |  |  |  |  |  | 29.00% | 41.00% |
Services Business | Discontinued Operations Held-for-sale | Nonrecurring Fair Value Measurements |  |  |  |  |  |  |  |  |
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
Impairment charges recorded |  |  | $ 1,350 |  | $ 1,350 |  |  |  |
Services Business | Discontinued Operations Disposed of by Sale |  |  |  |  |  |  |  |  |
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items] |  |  |  |  |  |  |  |  |
Impairment charges recorded |  |  |  |  |  | $ 0 | $ 1,350 | $ 0 |
Aggregate purchase price | $ 3,875 |  |  |  |  |  |  |  |
Working capital adjustment | 824 |  |  |  |  |  |  |  |
Consideration in escrow | $ 500 |  |  |  |  |  |  |  | $ 500
Consideration held in escrow, period held in escrow | 12 months |  |  |  |  |  |  |  |
Net cash consideration received at closing | $ 2,549 |  |  |  |  |  |  |  | $ 2,549

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