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Discontinued Operations - Additional Information (Details) - USD ($) $ in Thousands				1 Months Ended	3 Months Ended	12 Months Ended
	Jul. 21, 2015	Jul. 17, 2015	May 04, 2015	May 31, 2014	Jun. 30, 2015	Dec. 31, 2016	Dec. 31, 2015	Dec. 31, 2014	Jul. 16, 2015
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
Impairment charges recorded						$ 0	$ 586	$ 0
Consideration in escrow						0	500
Research and development						20,076	$ 16,440	$ 8,287
Continuing Operations
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
Research and development						$ 3,131
Research and Development Services
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
Contract initial term		18 months
Recorded unconditional purchase obligation		$ 3,300
Affiliate
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
General and administrative expense				$ 500
Investor | Merial | Sanofi
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
Ownership percentage of a related party in a customer of the company						100.00%
Investor | Customer Concentration Risk | Merial | Sales
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
Percentage of total revenue							29.00%	41.00%
Services Business | Discontinued Operations Held-for-sale | Nonrecurring Fair Value Measurements
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
Impairment charges recorded			$ 1,350		$ 1,350
Services Business | Discontinued Operations Disposed of by Sale
Income Statement, Balance Sheet and Additional Disclosures by Disposal Groups, Including Discontinued Operations [Line Items]
Impairment charges recorded						$ 0	$ 1,350	$ 0
Aggregate purchase price	$ 3,875
Working capital adjustment	824
Consideration in escrow	$ 500								$ 500
Consideration held in escrow, period held in escrow	12 months
Net cash consideration received at closing	$ 2,549								$ 2,549

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