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Stock Options - Stock Based Compensation Expense (Details) - USD ($) $ in Thousands | 12 Months Ended
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Dec. 31, 2021 | Dec. 31, 2020 | Dec. 31, 2019
Stock options |  |  |
Stock-based compensation expense included in Total operating expenses | $ 18,409 | $ 7,172 | $ 4,872
Stock options |  |  |
Stock options |  |  |
Stock-based compensation expense included in Total operating expenses | 14,528 | 5,725 | 4,230
Unvested restricted stock units |  |  |
Stock options |  |  |
Stock-based compensation expense included in Total operating expenses | 3,522 | 1,187 | 410
Employee stock purchase plan |  |  |
Stock options |  |  |
Stock-based compensation expense included in Total operating expenses | 359 | 260 | 232
Research and development |  |  |
Stock options |  |  |
Stock-based compensation expense included in Total operating expenses | 9,984 | 3,841 | 2,245
General and administrative |  |  |
Stock options |  |  |
Stock-based compensation expense included in Total operating expenses | $ 8,425 | $ 3,331 | $ 2,627

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