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Condensed Consolidated Statements of Stockholders’ Equity - USD ($) $ in Thousands | Total | At Market Equity Offering Program | Common Stock | Common Stock At Market Equity Offering Program | Additional Paid-in Capital | Additional Paid-in Capital At Market Equity Offering Program | Accumulated Other Comprehensive Income (Loss) | Accumulated Deficit
--- | --- | --- | --- | --- | --- | --- | --- | ---
Balance at beginning of period (in shares) at Dec. 31, 2022 |  |  | 105,144,864 |  |  |  |  |
Balance at beginning of period at Dec. 31, 2022 | $ 92,057 |  | $ 11 |  | $ 746,889 |  | $ (152) | $ (654,691)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Issuance of common stock from at-the-market transactions, net of issuance costs (in shares) |  |  |  | 3,535,093 |  |  |  |
Issuance of common stock from at-the-market transactions, net of issuance costs |  | $ 21,795 |  |  |  | $ 21,795 |  |
Exercise of stock options (in shares) |  |  | 8,826 |  |  |  |  |
Exercise of stock options | 34 |  |  |  | 34 |  |  |
Vesting of restricted stock units and other stock awards (in shares) |  |  | 372,291 |  |  |  |  |
Stock-based compensation expense | 6,407 |  |  |  | 6,407 |  |  |
Other comprehensive gain (loss) | 164 |  |  |  |  |  | 164 |
Net loss | (56,163) |  |  |  |  |  |  | (56,163)
Balance at end of period (in shares) at Mar. 31, 2023 |  |  | 109,061,074 |  |  |  |  |
Balance at end of period at Mar. 31, 2023 | 64,294 |  | $ 11 |  | 775,125 |  | 12 | (710,854)
Balance at beginning of period (in shares) at Dec. 31, 2022 |  |  | 105,144,864 |  |  |  |  |
Balance at beginning of period at Dec. 31, 2022 | 92,057 |  | $ 11 |  | 746,889 |  | (152) | (654,691)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Net loss | (152,126) |  |  |  |  |  |  |
Balance at end of period (in shares) at Sep. 30, 2023 |  |  | 120,548,980 |  |  |  |  |
Balance at end of period at Sep. 30, 2023 | 52,195 |  | $ 12 |  | 858,999 |  | 1 | (806,817)
Balance at beginning of period (in shares) at Mar. 31, 2023 |  |  | 109,061,074 |  |  |  |  |
Balance at beginning of period at Mar. 31, 2023 | 64,294 |  | $ 11 |  | 775,125 |  | 12 | (710,854)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Issuance of common stock from at-the-market transactions, net of issuance costs (in shares) |  |  |  | 10,929,438 |  |  |  |
Issuance of common stock from at-the-market transactions, net of issuance costs |  | 71,875 |  | $ 1 |  | 71,874 |  |
Exercise of stock options (in shares) |  |  | 88,770 |  |  |  |  |
Exercise of stock options | 393 |  |  |  | 393 |  |  |
Vesting of restricted stock units and other stock awards (in shares) |  |  | 88,690 |  |  |  |  |
Sales of common stock pursuant to ESPP (in shares) |  |  | 291,260 |  |  |  |  |
Sales of common stock pursuant to ESPP | 963 |  |  |  | 963 |  |  |
Stock-based compensation expense | 6,643 |  |  |  | 6,643 |  |  |
Other comprehensive gain (loss) | (61) |  |  |  |  |  | (61) |
Net loss | (54,307) |  |  |  |  |  |  | (54,307)
Balance at end of period (in shares) at Jun. 30, 2023 |  |  | 120,459,232 |  |  |  |  |
Balance at end of period at Jun. 30, 2023 | 89,800 |  | $ 12 |  | 854,998 |  | (49) | (765,161)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Vesting of restricted stock units and other stock awards (in shares) |  |  | 89,748 |  |  |  |  |
Stock-based compensation expense | 4,001 |  |  |  | 4,001 |  |  |
Other comprehensive gain (loss) | 50 |  |  |  |  |  | 50 |
Net loss | (41,656) |  |  |  |  |  |  | (41,656)
Balance at end of period (in shares) at Sep. 30, 2023 |  |  | 120,548,980 |  |  |  |  |
Balance at end of period at Sep. 30, 2023 | $ 52,195 |  | $ 12 |  | 858,999 |  | 1 | (806,817)
Balance at beginning of period (in shares) at Dec. 31, 2023 | 120,711,745 |  | 120,711,745 |  |  |  |  |
Balance at beginning of period at Dec. 31, 2023 | $ 36,904 |  | $ 12 |  | 863,242 |  | 11 | (826,361)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Issuance of common stock from at-the-market transactions, net of issuance costs (in shares) |  |  |  | 1,041,201 |  |  |  |
Issuance of common stock from at-the-market transactions, net of issuance costs |  | $ 5,780 |  |  |  | $ 5,780 |  |
Exercise of stock options (in shares) |  |  | 12,117 |  |  |  |  |
Exercise of stock options | 48 |  |  |  | 48 |  |  |
Vesting of restricted stock units and other stock awards (in shares) |  |  | 594,067 |  |  |  |  |
Stock-based compensation expense | 4,660 |  |  |  | 4,660 |  |  |
Other comprehensive gain (loss) | (101) |  |  |  |  |  | (101) |
Net loss | (19,306) |  |  |  |  |  |  | (19,306)
Balance at end of period (in shares) at Mar. 31, 2024 |  |  | 122,359,130 |  |  |  |  |
Balance at end of period at Mar. 31, 2024 | $ 27,985 |  | $ 12 |  | 873,730 |  | (90) | (845,667)
Balance at beginning of period (in shares) at Dec. 31, 2023 | 120,711,745 |  | 120,711,745 |  |  |  |  |
Balance at beginning of period at Dec. 31, 2023 | $ 36,904 |  | $ 12 |  | 863,242 |  | 11 | (826,361)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Exercise of stock options (in shares) | 14,223 |  |  |  |  |  |  |
Net loss | $ (55,075) |  |  |  |  |  |  |
Balance at end of period (in shares) at Sep. 30, 2024 | 122,866,659 |  | 122,866,659 |  |  |  |  |
Balance at end of period at Sep. 30, 2024 | $ 1,049 |  | $ 12 |  | 882,448 |  | 25 | (881,436)
Balance at beginning of period (in shares) at Mar. 31, 2024 |  |  | 122,359,130 |  |  |  |  |
Balance at beginning of period at Mar. 31, 2024 | 27,985 |  | $ 12 |  | 873,730 |  | (90) | (845,667)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Exercise of stock options (in shares) |  |  | 2,106 |  |  |  |  |
Exercise of stock options | 8 |  |  |  | 8 |  |  |
Vesting of restricted stock units and other stock awards (in shares) |  |  | 160,086 |  |  |  |  |
Sales of common stock pursuant to ESPP (in shares) |  |  | 153,729 |  |  |  |  |
Sales of common stock pursuant to ESPP | 258 |  |  |  | 258 |  |  |
Stock-based compensation expense | 4,425 |  |  |  | 4,425 |  |  |
Other comprehensive gain (loss) | 19 |  |  |  |  |  | 19 |
Net loss | (24,268) |  |  |  |  |  |  | (24,268)
Balance at end of period (in shares) at Jun. 30, 2024 |  |  | 122,675,051 |  |  |  |  |
Balance at end of period at Jun. 30, 2024 | 8,427 |  | $ 12 |  | 878,421 |  | (71) | (869,935)
Increase (Decrease) in Stockholders' Equity |  |  |  |  |  |  |  |
Vesting of restricted stock units and other stock awards (in shares) |  |  | 191,608 |  |  |  |  |
Stock-based compensation expense | 4,027 |  |  |  | 4,027 |  |  |
Other comprehensive gain (loss) | 96 |  |  |  |  |  | 96 |
Net loss | $ (11,501) |  |  |  |  |  |  | (11,501)
Balance at end of period (in shares) at Sep. 30, 2024 | 122,866,659 |  | 122,866,659 |  |  |  |  |
Balance at end of period at Sep. 30, 2024 | $ 1,049 |  | $ 12 |  | $ 882,448 |  | $ 25 | $ (881,436)

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