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Revenue - Collaboration Agreement - The GSK Termination and Transfer Agreement (Details) | 1 Months Ended | 3 Months Ended | 12 Months Ended |
--- | --- | --- | --- | ---
Sep. 30, 2023 GBP (£) | Jun. 30, 2023 GBP (£) | Mar. 31, 2024 USD ($) | Mar. 31, 2024 GBP (£) | Dec. 31, 2023 GBP (£) | Apr. 06, 2023 USD ($) | Apr. 06, 2023 GBP (£)
Revenue |  |  |  |  |  |  |
Revenue recognized in the period |  |  | $ 5,678,000 |  |  |  |
Aggregate amount of the transaction price that is allocated to performance obligations that are unsatisfied or partially satisfied under the agreements |  |  | 305,509,000 |  |  |  |
GSK Termination and Transfer Agreement |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |
Upfront payment received | £ |  | £ 7,500,000 |  |  |  |  |
Milestone payment | £ | £ 3,000,000 |  |  |  | £ 12,000,000 |  |
Potential milestone payments to be received | £ |  |  |  | £ 7,500,000 |  |  |
Milestone met and accrued | £ |  |  |  | £ 6,000,000 |  |  |
Amount of transaction price of the agreement at inception |  |  |  |  |  | $ 37,335,000 |
Upfront and milestone payment receivable | £ |  |  |  |  |  |  | £ 30,000,000
Aggregate amount of the transaction price that is allocated to performance obligations that are unsatisfied or partially satisfied under the agreements |  |  | 34,495,000 |  |  |  |
IGNYTE |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |
Aggregate amount of the transaction price that is allocated to performance obligations that are unsatisfied or partially satisfied under the agreements |  |  | 18,325,000 |  |  |  |
LTFU |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |
Aggregate amount of the transaction price that is allocated to performance obligations that are unsatisfied or partially satisfied under the agreements |  |  | $ 16,170,000 |  |  |  |

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