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Research Funding (Details) - USD ($) | 1 Months Ended | 12 Months Ended |
--- | --- | --- | ---
May 31, 2019 | Mar. 31, 2018 | Dec. 31, 2017 | May 31, 2017 | Dec. 31, 2019 | Dec. 31, 2018 | Feb. 28, 2017
Deferred Revenue Arrangement [Line Items] |  |  |  |  |  |  |
Grants receivable |  | $ 8,100,000 |  |  |  |  | $ 12,200,000
Funding of grant award |  |  |  | $ 5,700,000 |  |  |
Milestone payment received on grant award |  |  | $ 1,900,000 |  |  |  |
Progress payment received on grant award | $ 200,000 | $ 300,000 |  |  |  |  |
CIRM upfront funding - current |  |  |  |  | $ 1,600,000 | $ 2,583,000 |
Amortization of accrued grant funding to offset expense |  |  |  |  | 2,700,000 | $ 2,600,000 |
Accrued Liabilities |  |  |  |  |  |  |
Deferred Revenue Arrangement [Line Items] |  |  |  |  |  |  |
CIRM upfront funding - current |  |  |  |  | $ 1,600,000 |  |

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