<SEC-DOCUMENT>0001171843-21-003648.txt : 20210519
<SEC-HEADER>0001171843-21-003648.hdr.sgml : 20210519
<ACCEPTANCE-DATETIME>20210519170745
ACCESSION NUMBER:		0001171843-21-003648
CONFORMED SUBMISSION TYPE:	S-8
PUBLIC DOCUMENT COUNT:		5
FILED AS OF DATE:		20210519
DATE AS OF CHANGE:		20210519
EFFECTIVENESS DATE:		20210519

FILER:

	COMPANY DATA:	
		COMPANY CONFORMED NAME:			HARVARD BIOSCIENCE INC
		CENTRAL INDEX KEY:			0001123494
		STANDARD INDUSTRIAL CLASSIFICATION:	LABORATORY ANALYTICAL INSTRUMENTS [3826]
		IRS NUMBER:				043306140
		STATE OF INCORPORATION:			DE
		FISCAL YEAR END:			1231

	FILING VALUES:
		FORM TYPE:		S-8
		SEC ACT:		1933 Act
		SEC FILE NUMBER:	333-256295
		FILM NUMBER:		21941120

	BUSINESS ADDRESS:	
		STREET 1:		84 OCTOBER HILL RD
		CITY:			HOLLISTON
		STATE:			MA
		ZIP:			01746
		BUSINESS PHONE:		5088938999

	MAIL ADDRESS:	
		STREET 1:		84 OCTOBER HILL ROAD
		CITY:			HOLLISTON
		STATE:			MA
		ZIP:			01746
</SEC-HEADER>
<DOCUMENT>
<TYPE>S-8
<SEQUENCE>1
<FILENAME>s8_051821.htm
<DESCRIPTION>S-8
<TEXT>
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<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>As filed with the Securities and Exchange Commission on May 19, 2021</b></p>

<p style="font-size: 10pt; margin: 0pt 0"><b>&nbsp;</b></p>

<p style="font-size: 10pt; text-align: right; margin: 0pt 0"><b>Registration No. 333-_____</b></p>

<p style="font-size: 10pt; text-align: center; margin-top: 0pt; margin-bottom: 0pt"></p>

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<p style="font-size: 10pt; text-align: center; margin-top: 0pt; margin-bottom: 0pt"></p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>UNITED STATES</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>SECURITIES AND EXCHANGE COMMISSION</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>WASHINGTON, D.C. 20549</b></p>

<p style="font-size: 10pt; text-align: center; margin-top: 0pt; margin-bottom: 0pt"></p>

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<p style="font-size: 10pt; text-align: center; margin-top: 0pt; margin-bottom: 0pt">&nbsp;</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>FORM S-8<br>
REGISTRATION STATEMENT UNDER THE SECURITIES ACT OF 1933</b></p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt; text-align: center"></p>

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<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt; text-align: center"></p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>HARVARD BIOSCIENCE, INC.</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>(Exact Name of Registrant as Specified in Its Charter)</b></p>

<p style="font-size: 10pt; text-align: center; margin-top: 0pt; margin-bottom: 0pt"></p>

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<p style="font-size: 10pt; text-align: center; margin-top: 0pt; margin-bottom: 0pt">&nbsp;</p>

<table cellspacing="0" cellpadding="0" style="width: 100%; font-size: 10pt; border-collapse: collapse">
  <tr style="vertical-align: top">
    <td style="width: 49%; font-size: 10pt; text-align: center"><font style="font-size: 10pt"><b>Delaware<br>
(State of Incorporation)</b></font></td>
    <td style="width: 2%; font-size: 10pt">&nbsp;</td>
    <td style="width: 49%">
    <p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>04-3306140<br>
    (I.R.S. Employer</b></p>
    <p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>Identification No.)</b></p></td></tr>
  <tr style="vertical-align: top">
    <td style="font-size: 10pt">&nbsp;</td>
    <td style="font-size: 10pt">&nbsp;</td>
    <td style="font-size: 10pt">&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td>
    <p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>84 October Hill Road</b></p>
    <p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>Holliston, Massachusetts</b></p>
    <p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>(Address of Principal Executive Offices)</b></p></td>
    <td style="font-size: 10pt">&nbsp;</td>
    <td style="font-size: 10pt; text-align: center"><font style="font-size: 10pt"><b>01746<br>
(Zip Code)</b></font></td></tr>
  </table>
<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>HARVARD BIOSCIENCE, INC. 2021 INCENTIVE PLAN</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>(Full Title of the Plans)</b></p>

<p style="font-size: 10pt; text-align: justify; margin: 0pt 0"></p>

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<p style="margin: 0pt 0; font-size: 10pt; text-align: justify"></p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>James Green, Chief Executive Officer</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>HARVARD BIOSCIENCE, INC.</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>84 October Hill Road</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>Holliston, Massachusetts 01746</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>(508) 893-8999</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>(Name, address, and telephone number, including area code, of agent for
service)</b></p>

<p style="font-size: 10pt; text-align: justify; margin: 0pt 0"></p>

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<p style="margin: 0pt 0; font-size: 10pt; text-align: justify"></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b><i>&nbsp;</i></b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b><i>With copies to:</i></b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>Mitchell Raab</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>Olshan Frome Wolosky LLP</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>1325 Avenue of the Americas</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>New York, New York 10019</b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>(212) 451-2300</b></p>

<p style="font-size: 10pt; margin: 0pt 0"></p>

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<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>



<p style="margin: 0pt 0; font-size: 10pt">Indicate by check mark whether the registrant is a large accelerated filer, an accelerated filer,
a non-accelerated filer, a smaller reporting company, or an emerging growth company. See the definitions of &ldquo;large accelerated filer,&rdquo;
&ldquo;accelerated filer,&rdquo; &ldquo;smaller reporting company&rdquo; and &ldquo;emerging growth company&rdquo; in Rule 12b-2 of the
Exchange Act.</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<table cellpadding="0" cellspacing="0" style="border-collapse: collapse; width: 100%">
  <tr style="vertical-align: bottom">
    <td style="width: 50%; font-size: 10pt; text-align: left">Large accelerated filer &#9744;</td><td style="width: 1%; font-size: 10pt">&nbsp;</td>
    <td style="width: 49%; font-size: 10pt; text-align: left">Accelerated filer&#9746;</td></tr>
  <tr style="vertical-align: bottom">
    <td style="font-size: 10pt; text-align: left">Non-accelerated filer &#9744;</td><td style="font-size: 10pt">&nbsp;</td>
    <td style="font-size: 10pt; text-align: left">Smaller reporting company &#9746;</td></tr>
  <tr style="vertical-align: bottom">
    <td style="font-size: 10pt">&nbsp;</td><td style="font-size: 10pt">&nbsp;</td>
    <td style="font-size: 10pt; text-align: left">Emerging growth company&#9744;</td></tr>
  </table>


<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0">If an emerging growth company, indicate by check mark if the registrant has elected not to use
the extended transition period for complying with any new or revised financial accounting standards provided pursuant to Section 7(a)(2)(B)
of the Securities Act &#9744;</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

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<p style="margin: 0pt 0; font-size: 10pt"></p>

<table cellspacing="0" cellpadding="5" style="width: 100%; border-collapse: collapse; font-size: 10pt">
  <tr style="vertical-align: top">
    <td colspan="5" style="border: Black 1pt solid; text-align: center"><font style="font-size: 10pt"><b>Calculation of Registration Fee</b></font></td></tr>
  <tr style="vertical-align: bottom">
    <td style="border-left: Black 1pt solid; width: 20%; border-bottom: Black 1pt solid; border-right: Black 1pt solid; text-align: center"><font style="font-size: 10pt"><b>Title of Securities<br>
to be Registered</b></font></td>
    <td style="width: 20%; border-bottom: black 1pt solid; border-right: black 1pt solid; text-align: center"><font style="font-size: 10pt"><b>Amount<br>
to be<br>
Registered <sup>(1)</sup></b></font></td>
    <td style="width: 20%; border-bottom: black 1pt solid; border-right: black 1pt solid; text-align: center"><font style="font-size: 10pt"><b>Proposed<br>
Maximum<br>
Offering Price<br>
Per Share <sup>(2)</sup></b></font></td>
    <td style="width: 20%; border-bottom: black 1pt solid; border-right: black 1pt solid; text-align: center"><font style="font-size: 10pt"><b>Proposed<br>
Maximum<br>
Aggregate<br>
Offering Price <sup>(2)</sup></b></font></td>
    <td style="border-right: Black 1pt solid; border-bottom: Black 1pt solid; width: 20%; text-align: center"><font style="font-size: 10pt"><b>Amount of<br>
Registration Fee</b></font></td></tr>
  <tr style="vertical-align: top">
    <td style="border-left: Black 1pt solid; border-right: Black 1pt solid; border-bottom: Black 1pt solid"><font style="font-size: 10pt">Common Stock, par value $0.01 per share</font></td>
    <td style="border-right: black 1pt solid; border-bottom: Black 1pt solid; text-align: center"><font style="font-size: 10pt">4,293,756 shares <sup>(3)</sup></font></td>
    <td style="border-right: black 1pt solid; border-bottom: Black 1pt solid; text-align: center"><font style="font-size: 10pt">$6.92</font></td>
    <td style="border-right: black 1pt solid; border-bottom: Black 1pt solid; text-align: center"><font style="font-size: 10pt">$29,712,792</font></td>
    <td style="border-right: Black 1pt solid; border-bottom: Black 1pt solid; text-align: center"><font style="font-size: 10pt">$3,242</font></td></tr>
  </table>
<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0"></td><td style="width: 0.5in">(1)</td><td>Pursuant to Rule 416(a) under the Securities Act of 1933, as amended (the &ldquo;Securities Act&rdquo;), this Registration Statement
also covers such additional securities that become issuable by reason of any stock dividend, stock split, recapitalization or any other
similar transactions.</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0"></td><td style="width: 0.5in">(2)</td><td>Estimated solely for the purposes of calculating the registration fee pursuant to Rule 457(c) and 457(h) under the Securities Act,
based on the average of the high and low prices of the Common Stock as reported on the Nasdaq Global Market on May 13, 2021.&#9;</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0"></td><td style="width: 0.5in">(3)</td><td>Consists of shares of common stock, par value $0.01 per share (&ldquo;Common Stock&rdquo;), of Harvard Bioscience, Inc. (the &ldquo;Registrant&rdquo;)
that are issuable pursuant to the Registrant&rsquo;s 2021 Incentive Plan.</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center"></p>

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<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">PART I<br>
INFORMATION REQUIRED IN THE SECTION 10(a) PROSPECTUS</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">The information called for in Part I of Form S-8 to be contained in the
Section 10(a) prospectus is not being filed with or included in this registration statement (by incorporation by reference or otherwise)
in accordance with the rules and regulations of the Securities and Exchange Commission (the &ldquo;Commission&rdquo;).</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0">PART II<br>
INFORMATION REQUIRED IN THE REGISTRATION STATEMENT</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">Item 3. Incorporation of Documents by Reference.</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">The Registrant is subject to the informational and reporting requirements
of Sections 13(a), 14 and 15(d) of the Securities Exchange Act of 1934, as amended (the &ldquo;Exchange Act&rdquo;), and in accordance
therewith files reports, proxy statements and other information with the Commission. The following documents, which are on file with the
Commission, are hereby incorporated by reference in, and shall be deemed a part of, this Registration Statement (other than any such documents
or portions thereof that are furnished under Item 2.02 or Item 7.01 of Form 8-K, unless otherwise indicated therein, including any exhibits
included with such Items):</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">(a)&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;The Registrant&rsquo;s Annual
Report on Form 10-K for the year ended December 31, 2020 filed with the Commission on <a href="https://www.sec.gov/ix?doc=/Archives/edgar/data/1123494/000117184321001737/hboi20201231_10k.htm">March
12, 2021</a>, as amended by the Annual Report on Form 10-K/A for the year ended December 31, 2020 filed with the Commission on <a href="https://www.sec.gov/ix?doc=/Archives/edgar/data/1123494/000117184321001848/hboi20201231_10ka.htm">March
17, 2021</a>;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">(b)&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;The Registrant&rsquo;s Definitive
Proxy Statement on Schedule 14A filed with the Commission on <a href="https://www.sec.gov/Archives/edgar/data/1123494/000117184321002394/def14a_040521.htm">April
8, 2021</a>;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">(c)&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;The Registrant&rsquo;s Quarterly
Report on Form 10-Q for the quarter ended March 31, 2021 filed with the Commission on <a href="https://www.sec.gov/ix?doc=/Archives/edgar/data/1123494/000117184321003314/hboi20210331_10q.htm">May
7, 2021</a>;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">(d)&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;The Registrant&rsquo;s Current
Report on Form 8-K filed with the Commission on <a href="https://www.sec.gov/Archives/edgar/data/1123494/000117184321003642/f8k_051821.htm">May 19, 2021</a>; and</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">(e)&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;The description of the Registrant&rsquo;s
Common Stock contained in the Registrant&rsquo;s registration statement on Form 8-A (File No. 000-31923) filed with the Commission on
<a href="https://www.sec.gov/Archives/edgar/data/1123494/000091205700047956/a2029213z8-a12g.txt">November 9, 2000</a> under the Exchange
Act, including any amendment or report filed for the purpose of updating such description.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">All documents subsequently filed by the Registrant pursuant to Sections
13(a), 13(c), 14 and 15(d) of the Exchange Act, prior to the filing of a post-effective amendment which indicates that all securities
offered hereby have been sold or which deregisters all securities then remaining unsold, shall be deemed to be incorporated by reference
in this Registration Statement and to be part hereof from the date of the filing of such documents. The Registrant is not, however, incorporating
by reference any documents or portions thereof, whether specifically listed above or filed in the future, that are not deemed &ldquo;filed&rdquo;
with the Commission, including any information furnished pursuant to Items 2.02 or 7.01 of Form 8-K or related exhibits furnished pursuant
to Item 9.01 of Form 8-K.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-indent: 0.5in">&nbsp;</p>

<p style="text-align: left; margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0"></p>

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<p style="margin: 0pt 0; font-size: 10pt; text-indent: 0.5in">Any statement contained in a document incorporated or deemed to be incorporated
by reference herein shall be deemed to be modified or superseded for the purposes of this Registration Statement to the extent that a
statement contained herein or in any other subsequently filed document which also is or is deemed to be incorporated by reference herein
modifies or supersedes such statement. Any statement so modified or superseded shall not be deemed, except as so modified or superseded,
to constitute a part of this Registration Statement. Under no circumstances will any information furnished under Items 2.02 or 7.01 of
Form 8-K be deemed incorporated by reference unless such Current Report on Form 8-K expressly provides to the contrary.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">Item 4. Description of Securities.</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">Not applicable.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">Item 5. Interests of Named Experts and Counsel.</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">Not applicable.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">Item 6. Indemnification of Directors and Officers.</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">As permitted by Section 102 of the Delaware General Corporation Law, the
Registrant has adopted provisions in its Second Amended and Restated Certificate of Incorporation, or its Charter, and Amended and Restated
Bylaws, as amended, or its Bylaws, that limit or eliminate the personal liability of its directors for a breach of their fiduciary duty
of care as a director. The duty of care generally requires that, when acting on behalf of a corporation, directors exercise an informed
business judgment based on all material information reasonably available to them. Consequently, a director will not be personally liable
to the Registrant or its stockholders for monetary damages for breach of fiduciary duty as a director, except for liability for:</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.5in"></td><td style="width: 0.5in"><font style="font-family: Symbol">&middot;</font></td><td>any breach of the director&rsquo;s duty of loyalty to the Registrant or its stockholders;</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.5in"></td><td style="width: 0.5in"><font style="font-family: Symbol">&middot;</font></td><td>any act or omission not in good faith or that involves intentional misconduct or a knowing violation of law;</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.5in"></td><td style="width: 0.5in"><font style="font-family: Symbol">&middot;</font></td><td>any act related to unlawful stock repurchases, redemptions or other distributions or payments of dividends; or</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.5in"></td><td style="width: 0.5in"><font style="font-family: Symbol">&middot;</font></td><td>any transaction from which the director derived an improper personal benefit.</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">These limitations of liability do not affect the availability of equitable
remedies such as injunctive relief or rescission. The Registrant&rsquo;s Bylaws also authorizes the Registrant to indemnify its officers,
directors and other agents to the fullest extent permitted under Delaware law.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0"></p>

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<p style="margin: 0pt 0; font-size: 10pt; text-indent: 0.5in">As permitted by Section 145 of the Delaware General Corporation Law, the
Registrant&rsquo;s Bylaws provide that it will indemnify any person who was or is a party or is threatened to be made a party to any threatened,
pending or completed action, suit or other proceeding by reason of the fact that such person, or a person of whom he or she is the legal
representative, is or was the Registrant&rsquo;s director or officer, or by reason of the fact that the Registrant&rsquo;s director or
officer is or was serving, at the Registrant&rsquo;s request, as a director, partner, trustee, officer, employee or agent of another corporation
or of a partnership, joint venture, trust or other enterprise, including service with respect to employee benefit plans maintained or
sponsored by the Registrant. The Registrant will indemnify such persons against expenses (including reasonable attorneys&rsquo; fees),
judgments, penalties, fines and amounts reasonably paid in settlement that are incurred in connection with such action if such person
acted in good faith and in a manner reasonably believed to be in or not opposed to the Registrant&rsquo;s best interests and, with respect
to any criminal proceeding, had no reasonable cause to believe such person&rsquo;s conduct was unlawful. Any amendment of this provision
will not reduce the Registrant&rsquo;s indemnification obligations relating to actions taken before an amendment.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">In addition, the Registrant has entered into separate indemnification agreements,
a form of which is attached as Exhibit 10.2 to the Registrant&rsquo;s Quarterly Report on Form 10-Q (filed with the Commission on May
8, 2020), with the Registrant&rsquo;s directors and certain of its officers which may be broader than the specific indemnification provisions
contained in the Delaware General Corporation Law. These indemnification agreements generally require the Registrant, among other things,
to indemnify such officers and directors against liabilities that may arise by reason of their status or service as directors or officers,
subject to certain exceptions and limitations. These indemnification agreements also require the Registrant to advance any expenses incurred
by such directors or officers as a result of any proceeding against them as to which they could be indemnified.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">The Registrant has also obtained policies that insure its directors and
officers and those of its subsidiaries against certain liabilities they may incur in their capacity as directors and officers. Under these
policies, the insurer, on the Registrant&rsquo;s behalf, may also pay amounts for which the Registrant has granted indemnification to
the directors or officers. These indemnification provisions and the indemnification agreements may be sufficiently broad to permit indemnification
of the Registrant&rsquo;s officers and directors for liabilities, including reimbursement of expenses incurred, arising under the Securities
Act.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">The foregoing summaries are qualified in their entirety by the terms and
provisions of such arrangements.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">Item 7. Exemption from Registration Claimed.</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">Not applicable.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">Item 8. Exhibits.</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">The Exhibit Index preceding the signature page to this Registration Statement
is incorporated by reference.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0"></p>

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<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold">Item 9. Undertakings.</p>

<p style="font-size: 10pt; font-weight: bold; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0">(a) The undersigned registrant hereby undertakes:</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.25in"></td><td style="width: 0.25in">(1)</td><td>To file, during any period in which offers or sales are being made, a post-effective amendment to this Registration Statement:</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.5in"></td><td style="width: 0.5in">(i)</td><td>to include any prospectus required by Section 10(a)(3) of the Securities Act;</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.5in"></td><td style="width: 0.5in">(ii)</td><td>to reflect in the prospectus any facts or events arising after the effective date of the Registration Statement (or the most recent
post-effective amendment thereof) which, individually or in the aggregate, represent a fundamental change in the information set forth
in the Registration Statement. Notwithstanding the foregoing, any increase or decrease in volume of securities offered (if the total dollar
value of securities offered would not exceed that which was registered) and any deviation from the low or high end of the estimated maximum
offering range may be reflected in the form of prospectus filed with the Commission pursuant to Rule 424(b) if, in the aggregate, the
changes in volume and price represent no more than 20 percent change in the maximum aggregate offering price set forth in the &ldquo;Calculation
of Registration Fee&rdquo; table in the effective Registration Statement; and</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.5in"></td><td style="width: 0.5in">(iii)</td><td>to include any material information with respect to the plan of distribution not previously disclosed in the Registration Statement
or any material change to such information in the Registration Statement;</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0 0pt 0.5in"><i>provided, however</i>, that paragraphs (a)(1)(i) and (a)(1)(ii) do not apply if
the information required to be included in a post-effective amendment by those paragraphs is contained in reports filed with or furnished
to the Commission by the registrant pursuant to Section 13 or Section 15(d) of the Exchange Act that are incorporated by reference in
the Registration Statement.</p>

<p style="font-size: 10pt; margin: 0pt 0 0pt 0.5in">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.25in"></td><td style="width: 0.25in">(2)</td><td>That, for the purpose of determining any liability under the Securities Act, each such post-effective amendment shall be deemed to
be a new registration statement relating to the securities offered therein, and the offering of such securities at that time shall be
deemed to be the initial bona fide offering thereof.</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0.25in"></td><td style="width: 0.25in">(3)</td><td>To remove from registration by means of a post-effective amendment any of the securities being registered which remain unsold at the
termination of the offering.</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0">(b) The undersigned registrant hereby undertakes that, for purposes of determining any liability
under the Securities Act, each filing of the registrant&rsquo;s annual report pursuant to Section 13(a) or 15(d) of the Exchange Act (and,
where applicable, each filing of an employee benefit plan&rsquo;s annual report pursuant to Section 15(d) of the Exchange Act) that is
incorporated by reference in this Registration Statement shall be deemed to be a new registration statement relating to the securities
offered therein, and the offering of such securities at that time shall be deemed to be the initial bona fide offering thereof.</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>



<p style="margin: 0pt 0; font-size: 10pt">(h) Insofar as indemnification for liabilities arising under the Securities Act may be permitted
directors, officers and controlling persons of the registrant pursuant to the foregoing provisions, or otherwise, the registrant has been
advised that in the opinion of the Commission such indemnification is against public policy as expressed in the Securities Act and is,
therefore, unenforceable. In the event that a claim for indemnification against such liabilities (other than the payment by the registrant
of expenses incurred or paid by a director, officer or controlling person of the registrant in the successful defense of any action, suit
or proceeding) is asserted by such director, officer or controlling person in connection with the securities being registered, the registrant
will, unless in the opinion of its counsel the matter has been settled by controlling precedent, submit to a court of appropriate jurisdiction
the question whether such indemnification by it is against public policy as expressed in the Securities Act and will be governed by the
final adjudication of such issue.</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center"></p>

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    <div style="break-before: page; margin-top: 6pt; margin-bottom: 12pt"><p style="margin: 0pt">&#160;</p></div>
    <!-- Field: /Page -->

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">EXHIBIT INDEX</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0">&nbsp;</p>

<table cellspacing="0" cellpadding="5" style="width: 100%; border-collapse: collapse; font-size: 10pt">
  <tr style="vertical-align: bottom">
    <td style="width: 12%">
    <p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0; border-bottom: Black 0.5pt solid"><b>Exhibit<br>
Number</b></p></td>
    <td style="width: 88%">
    <p style="font-size: 10pt; text-align: center; text-indent: 0in; margin: 0pt 0; border-bottom: Black 0.5pt solid"><b>Description</b></p></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">4.1<sup>(1)</sup></td>
    <td style="text-indent: 0in"><a href="https://www.sec.gov/Archives/edgar/data/1123494/000091205700047957/a2029062zex-3_2.txt">Second Amended and Restated Certificate of Incorporation of Harvard Bioscience, Inc.</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">4.2<sup>(1)</sup></td>
    <td style="text-indent: 0in"><a href="https://www.sec.gov/Archives/edgar/data/1123494/000091205700047957/a2029062zex-3_3.txt">Amended and Restated Bylaws of Harvard Bioscience, Inc.</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">4.3<sup>(2)</sup></td>
    <td style="text-indent: 0in"><a href="https://www.sec.gov/Archives/edgar/data/1123494/000119312507232564/dex31.htm">Amendment No. 1 to Amended and Restated Bylaws of Harvard Bioscience, Inc.</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">4.4<sup>(1)</sup></td>
    <td style="text-indent: 0in"><a href="https://www.sec.gov/Archives/edgar/data/1123494/000091205700047957/a2029062zex-4_1.txt">Specimen certificate for shares of Common Stock, $0.01 par value, of Harvard Bioscience, Inc.</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">4.5<sup>(3)</sup></td>
    <td style="text-indent: 0in"><a href="https://www.sec.gov/Archives/edgar/data/1123494/000117184321003642/exh_101.htm">Harvard Bioscience, Inc. 2021 Incentive Plan.</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">5.1*</td>
    <td style="text-indent: 0in"><a href="exh_51.htm">Opinion of Olshan Frome Wolosky LLP.</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">23.1*</td>
    <td style="text-indent: 0in"><a href="exh_231.htm">Consent of Grant Thornton LLP.</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">23.2*</td>
    <td style="text-indent: 0in"><a href="exh_51.htm">Consent of Olshan Frome Wolosky LLP (contained in Exhibit 5.1).</a></td></tr>
  <tr style="vertical-align: top">
    <td style="text-indent: 0in">24.1*</td>
    <td style="text-indent: 0in"><a href="#a_001">Power of attorney (included on the signature page to this Registration Statement).</a></td></tr>
  </table>

<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0"></td><td style="width: 0.25in">*</td><td>Filed herewith.</td></tr></table>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0"></td><td style="width: 0.25in"><sup>(1)</sup></td><td>Previously filed as an exhibit to the Company&rsquo;s Registration Statement on Form S-1/A (File No. 333-45996) (filed on November
9, 2000) and incorporated by reference thereto.</td></tr></table>

<p style="margin-top: 0; margin-bottom: 0">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0"></td><td style="width: 0.25in"><sup>(2)</sup></td><td>Previously filed as an exhibit to the Company&rsquo;s Current Report on Form 8-K (filed on November 1, 2007) and incorporated by reference
thereto.</td></tr></table>

<p style="margin-top: 0; margin-bottom: 0">&nbsp;</p>

<table cellpadding="0" cellspacing="0" width="100%" style="font-size: 10pt; margin-top: 0pt; margin-bottom: 0pt"><tr style="vertical-align: top">
<td style="width: 0"></td><td style="width: 0.25in"><sup>(3)</sup></td><td>Previously filed as an exhibit to the Company&rsquo;s Current Report on Form 8-K (filed on May 19, 2021) and incorporated by reference
thereto.</td></tr></table>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center"></p>

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    <div style="break-before: page; margin-top: 6pt; margin-bottom: 12pt"><p style="margin: 0pt">&#160;</p></div>
    <!-- Field: /Page -->

<p style="margin: 0pt 0; font-size: 10pt; font-weight: bold; text-align: center">SIGNATURES</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">Pursuant to the requirements of the Securities Act of 1933, the Registrant
certifies that it has reasonable grounds to believe that it meets all of the requirements for filing on Form S-8 and has duly caused this
registration statement to be signed on its behalf by the undersigned, thereunto duly authorized, in the Town of Holliston, Massachusetts,
on this 19<sup>th</sup> day of May, 2021.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<table cellspacing="0" cellpadding="0" style="width: 100%; border-collapse: collapse; font-size: 10pt">
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td colspan="2">HARVARD BIOSCIENCE, INC.</td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td colspan="2">&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td style="width: 51%">&nbsp;</td>
    <td style="width: 6%">By:</td>
    <td style="border-bottom: Black 1pt solid; width: 43%">/s/ James Green</td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td>&nbsp;</td>
    <td><i>James Green, Chief Executive Officer</i></td></tr>
  </table>
<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0"><a name="a_001"></a>POWER OF ATTORNEY</p>

<p style="font-size: 10pt; font-weight: bold; text-align: center; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">KNOW ALL BY THESE PRESENTS that each individual whose signature appears
below constitutes and appoints each of James Green and Michael A. Rossi as such person&rsquo;s true and lawful attorney-in-fact and agent
with full power of substitution and resubstitution, for such person in such person&rsquo;s name, place and stead, in any and all capacities,
to sign any and all amendments (including post-effective amendments) to this registration statement (or any registration statement for
the same offering that is to be effective upon filing pursuant to Rule 462(b) under the Securities Act of 1933), and to file the same,
with all exhibits thereto, and all documents in connection therewith, with the Securities and Exchange Commission, granting unto each
said attorney-in-fact and agent full power and authority to do and perform each and every act and thing requisite and necessary to be
done in and about the premises, as fully to all intents and purposes as such person might or could do in person, hereby ratifying and
confirming all that any said attorney-in-fact and agent, or any substitute or substitutes of any of them, may lawfully do or cause to
be done by virtue hereof.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">Pursuant to the requirements of the Securities Act of 1933, this registration
statement has been signed by the following persons in the capacities and on the date indicated. Each person listed below has signed this
registration statement as an officer or director of Harvard Bioscience, Inc.</p>

<p style="font-size: 10pt; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<table cellspacing="0" cellpadding="3" style="width: 100%; border-collapse: collapse; font-size: 10pt">
  <tr style="vertical-align: bottom">
    <td style="width: 33%; text-align: center"><b>Signature</b></td>
    <td style="width: 46%; text-align: center"><b>Title</b></td>
    <td style="width: 21%; text-align: center"><b>Date</b></td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td style="border-bottom: Black 1pt solid">/s/ James Green</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td>James Green</td>
    <td><p style="margin-top: 0; margin-bottom: 0">Chief Executive Officer and Director</p>
        <p style="margin-top: 0; margin-bottom: 0">(Principal Executive Officer)</p></td>
    <td>May 19, 2021</td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td style="border-bottom: Black 1pt solid">/s/ Michael A. Rossi</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td>Michael A. Rossi</td>
    <td>Chief Financial Officer</td>
    <td>May 19, 2021</td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td style="border-bottom: Black 1pt solid">/s/ Katherine A. Eade</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td>Katherine A. Eade</td>
    <td>Director</td>
    <td>May 19, 2021</td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td style="border-bottom: Black 1pt solid">/s/ Alan Edrick</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td>Alan Edrick</td>
    <td>Director</td>
    <td>May 19, 2021</td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td style="border-bottom: Black 1pt solid">/s/ Thomas W. Loewald</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td>Thomas W. Loewald</td>
    <td>Director</td>
    <td>May 19, 2021</td></tr>
  <tr style="vertical-align: top">
    <td>&nbsp;</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td style="border-bottom: Black 1pt solid">/s/ Bertrand Loy</td>
    <td>&nbsp;</td>
    <td>&nbsp;</td></tr>
  <tr style="vertical-align: top">
    <td>Bertrand Loy</td>
    <td>Director</td>
    <td>May 19, 2021</td></tr>
  </table>


<p style="margin: 0">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>

<p style="text-align: center; margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt">6</p>

<!-- Field: Rule-Page --><div style="margin: 3pt auto; width: 100%"><div style="font-size: 1pt; border-top: Black 4pt solid">&nbsp;</div></div><!-- Field: /Rule-Page -->

<p style="text-align: center; margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt"></p>

<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>

</body>
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</DOCUMENT>
<DOCUMENT>
<TYPE>EX-5.1
<SEQUENCE>2
<FILENAME>exh_51.htm
<DESCRIPTION>EXHIBIT 5.1
<TEXT>
<html>
<head>
     <title></title>
</head>
<body style="font: 10pt Times New Roman, Times, Serif">

<p style="text-align: right; margin: 0"><b>Exhibit 5.1</b></p>

<p style="margin: 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: center"><img src="exh51hdr.jpg" alt=""></p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: center">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: center">May 19, 2021</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0">Harvard Bioscience, Inc.</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0">84 October Hill Road</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0">Holliston, Massachusetts 01746</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 1in; margin: 0pt 0">Re: <u>Registration Statement on Form S-8</u></p>

<p style="font-size: 10pt; text-indent: 1in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0">Ladies and Gentlemen:</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">We have acted as counsel to Harvard Bioscience, Inc.,
a Delaware corporation (the &ldquo;Company&rdquo;), in connection with the filing with the Securities and Exchange Commission (the &ldquo;Commission&rdquo;)
of the Company&rsquo;s Registration Statement on Form S-8 (the &ldquo;Registration Statement&rdquo;) relating to the registration of 4,293,756
shares (the &ldquo;Shares&rdquo;) of the Company&rsquo;s common stock, par value $0.01 per share (the &ldquo;Common Stock&rdquo;), issuable
pursuant to the terms of and in the manner set forth in the Company&rsquo;s 2021 Incentive Plan (the &ldquo;Plan&rdquo;). This opinion
letter is being delivered at the request of the Company and in accordance with the requirements of Item 601(b)(5) of Regulation S-K promulgated
under the Securities Act of 1933, as amended (the &ldquo;Securities Act&rdquo;).</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">We advise you that we have examined executed originals
or copies certified or otherwise identified to our satisfaction of (i) the Registration Statement, (ii) the Company&rsquo;s Second Amended
and Restated Certificate of Incorporation and Amended and Restated Bylaws, each as amended to date, (iii) the Plan, and (iv) corporate
proceedings of the Company, and such other documents, instruments and certificates of officers and representatives of the Company and
of public officials, and we have made such examination of law, as we have deemed necessary or appropriate for purposes of the opinion
expressed below.</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">We have assumed for purposes of rendering the opinion
set forth herein, without any verification by us, the genuineness of all signatures, the legal capacity of all natural persons to execute
and deliver documents, the authenticity and completeness of documents submitted to us as originals and the completeness and conformity
with authentic original documents of all documents submitted to us as copies, and that all documents, books and records made available
to us by the Company are accurate and complete.</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">On the basis of the foregoing and in reliance thereon
and subject to the assumptions, qualifications and limitations set forth herein, we advise you that in our opinion, the Shares, when issued
and paid for pursuant to the terms of and in the manner set forth in the Plan, will be duly and validly issued, fully paid and non-assessable.</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: justify; text-indent: 0.5in">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: justify; text-indent: 0.5in">&nbsp;</p>

<p style="margin-top: 0pt; margin-bottom: 0pt; font-size: 10pt; text-align: center"><img src="exh51foot.jpg" alt=""></p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0"></p>

<!-- Field: Page; Sequence: 1 -->
    <div style="margin-top: 12pt; margin-bottom: 6pt; border-bottom: Black 4pt solid"><p style="margin: 0pt">&#160;</p></div>
    <div style="break-before: page; margin-top: 6pt; margin-bottom: 12pt"><p style="margin: 0pt">&#160;</p></div>
    <!-- Field: /Page -->

<p style="font-size: 10pt; margin: 0pt 0">May 19, 2021</p>

<p style="font-size: 10pt; margin: 0pt 0">Page 2</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: justify; text-indent: 0.5in">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: justify; text-indent: 0.5in">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt; text-align: justify; text-indent: 0.5in">We are members of the Bar of the State of New York.
We express no opinion as to the effect of any laws other than the laws of the State of New York, the General Corporation Law of the State
of Delaware and the Federal laws of the United States of America, each as in effect on the date hereof.</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">This opinion speaks only at and as of its date and
is based solely on the facts and circumstances known to us at and as of such date. We assume no obligation to revise or supplement this
opinion to reflect any facts or circumstances that may hereafter come to our attention or any changes in fact or law that may hereafter
occur.</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">We hereby consent to the filing of this opinion as
an exhibit to the Registration Statement. In giving such consent, we do not thereby concede that our firm is within the category of persons
whose consent is required under Section 7 of the Securities Act or the rules and regulations of the Commission thereunder.</p>

<p style="font-size: 10pt; text-align: justify; text-indent: 0.5in; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0 0pt 3in">Very truly yours,</p>

<p style="font-size: 10pt; margin: 0pt 0 0pt 3in">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0 0pt 3in">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0 0pt 3in">&nbsp;</p>

<p style="margin: 0pt 0 0pt 3in; font-size: 10pt">/s/ Olshan Frome Wolosky LLP</p>

<p style="margin: 0pt 0 0pt 3in; font-size: 10pt">&nbsp;</p>

<p style="margin: 0pt 0 0pt 3in; font-size: 10pt">OLSHAN FROME WOLOSKY LLP</p>

<p style="font-size: 10pt; text-indent: 0in; margin: 0pt 0 0pt 3in">&nbsp;</p>
<p style="margin: 0">&#160;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

<p style="margin: 0">&nbsp;</p>

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</TEXT>
</DOCUMENT>
<DOCUMENT>
<TYPE>EX-23.1
<SEQUENCE>3
<FILENAME>exh_231.htm
<DESCRIPTION>EXHIBIT 23.1
<TEXT>
<html>
<head>
     <title></title>
</head>
<body style="font: 10pt Times New Roman, Times, Serif">

<p style="text-align: right; margin: 0"><b>Exhibit 23.1</b></p>

<p style="margin: 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>CONSENT OF INDEPENDENT REGISTERED PUBLIC ACCOUNTING FIRM </b></p>

<p style="font-size: 10pt; text-align: center; margin: 0pt 0"><b>&nbsp;</b></p>

<p style="font-size: 10pt; margin: 0pt 0">We have issued our reports dated March 12, 2021, with respect to the consolidated financial
statements and internal control over financial reporting of Harvard Bioscience, Inc., included in the Annual Report on Form 10-K for the
year ended December 31, 2020, which are incorporated by reference in this Registration Statement. We consent to the incorporation by reference
of the aforementioned reports in this Registration Statement.</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; text-align: justify; margin: 0pt 0">/s/ GRANT THORNTON LLP</p>

<p style="font-size: 10pt; text-align: justify; margin: 0pt 0">&nbsp;</p>

<p style="font-size: 10pt; margin: 0pt 0">Boston, Massachusetts</p>

<p style="font-size: 10pt; margin: 0pt 0">May 19, 2021</p>

<p style="font-size: 10pt; margin: 0pt 0">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>

<p style="margin: 0pt 0; font-size: 10pt">&nbsp;</p>

</body>
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end
</TEXT>
</DOCUMENT>
</SEC-DOCUMENT>
