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Consolidated Statements of Cash Flows (Unaudited) - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Cash flows from operating activities:    
Net loss $ (15,697) $ (14,971)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation 601 626
Equity-based compensation expense 890 877
Accretion/amortization of investment securities (11) 105
Change in carrying amount of operating lease right of use asset 769 462
Changes in operating assets and liabilities:    
Prepaid expenses and other current assets 821 (443)
Prepaid research and development, net of current portion (488) 1,382
Accounts payable and accrued expenses (1,387) (1,408)
Deferred revenue (217)  
Operating lease liabilities (739) (606)
Other assets (19)  
Net cash used in operating activities (15,477) (13,976)
Cash flows from investing activities:    
Purchases of marketable securities (12,121) (27,091)
Proceeds from maturity of marketable securities 32,356 25,996
Proceeds from redemption of marketable securities   1,270
Purchases of property and equipment (477) (21)
Net cash provided by investing activities 19,758 154
Cash flows from financing activities:    
Payments on finance lease obligations (3) (1)
Proceeds from employee stock purchases and exercise of stock options 135 33
Payment of employee withholding taxes relating to restricted stock awards   (65)
Net cash provided by financing activities 132 8,019
Net increase (decrease) in cash, cash equivalents and restricted cash 4,413 (5,803)
Cash, cash equivalents and restricted cash at beginning of period 17,535 33,604
Cash, cash equivalents and restricted cash at end of period 21,948 27,801
Supplemental disclosure of non-cash investing activities:    
Assets acquired under operating lease obligation 1,773  
Property and equipment purchases included in accounts payable and accrued expenses 118 3
Supplemental disclosure of non-cash financing activities:    
Issuance costs included in accounts payable and accrued expenses   2
Cash paid for interest $ 1  
At-the-market Offering    
Cash flows from financing activities:    
Proceeds from issuance of common stock, net of issuance costs   $ 8,052