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Stock-Based Compensation - Stock options (Details) - USD ($) $ / shares in Units, $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2020 | Dec. 31, 2019 | Dec. 31, 2018
Options |  |  |
Outstanding balance at the beginning of the period (in shares) | 4,024,566 | 2,761,912 |
Granted (in shares) | 2,662,500 | 1,481,975 |
Exercised (in shares) | (32,778) |  |
Forfeited (in shares) | (260,435) | (219,321) |
Outstanding balance at the end of the period (in shares) | 6,393,853 | 4,024,566 | 2,761,912
Options exercisable at the end of the period (in shares) | 3,482,169 |  |
Options vested and expected to vest at end of period (in shares) | 6,393,853 |  |
Weighted Average Exercise Price Per Share |  |  |
Outstanding balance at the beginning of the period (in dollars per share) | $ 7.48 | $ 11.30 |
Granted (in dollars per share) | 2.10 | 1.48 |
Exercised (in dollars per share) | 1.53 |  |
Forfeited (in dollars per share) | 6.95 | 15.14 |
Outstanding balance at the end of the period (in dollars per share) | 5.29 | $ 7.48 | $ 11.30
Exercisable at the end of the period (in dollars per share) | 7.83 |  |
Options vested and expected to vest at end of the period (in dollars per share) | $ 5.29 |  |
Weighted Average Remaining Life in Years |  |  |
Outstanding balance | 7 years 11 months 16 days | 7 years 11 months 23 days | 8 years 4 days
Options exercisable at end of period | 7 years 26 days |  |
Options vested and expected to vest at end of the period | 7 years 11 months 16 days |  |
Aggregate Intrinsic Value |  |  |
Outstanding balance | $ 1,961 | $ 2 |
Options exercisable at the end of the period | 722 |  |
Options vested and expected to vest at end of the period | 1,961 |  |
Stock-based compensation expense | 2,572 | 3,732 |
Stock-based compensation | $ 2,324 | $ 3,732 |
Assumptions used to determine grant-date fair value |  |  |
Expected Volatility | 66.94% | 66.94% |
Weighted average risk-free interest rate | 0.66% | 2.07% |
Expected dividend yield (as a percentage) | 0.00% | 0.00% |
Expected term (in years) | 6 years 10 months 6 days | 6 years 9 months 7 days |
Expiration period | 10 years |  |
Unrecognized costs | $ 3,947 |  |
Research and development |  |  |
Aggregate Intrinsic Value |  |  |
Stock-based compensation expense | 1,028 | $ 714 |
General and administrative |  |  |
Aggregate Intrinsic Value |  |  |
Stock-based compensation expense | $ 1,296 | $ 3,018 |
Weighted average |  |  |
Assumptions used to determine grant-date fair value |  |  |
Weighted average period for recognition of compensation cost | 2 years 29 days |  |
Minimum |  |  |
Assumptions used to determine grant-date fair value |  |  |
Vesting period | 3 years |  |
Maximum |  |  |
Assumptions used to determine grant-date fair value |  |  |
Vesting period | 4 years |  |

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