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Equity Incentive Plans (Tables)

9 Months Ended

Sep. 30, 2024

Equity Incentive Plans

Schedule of stock option activity under the Equity Plans

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​	​	​	​	​	​	​	​	​	​	​
​		​	​		Weighted		​	​		​
​	​	​	​		Average		Weighted	​		Aggregate
​	​	​	​		Exercise Price		Average Remaining	​		Intrinsic
​		Options	​		Per Share		Life in Years	​		Value
Outstanding at December 31, 2023		3,384,366	​	$	13.97		8.40	​	$	3,431
Granted	​	1,727,500	​	$	2.67	​	​	​	​	​
Exercised	​	(10,557)	​	$	2.77	​	​	​	​	​
Forfeited		(254,674)	​	$	5.69		​	​	​	​
Outstanding at September 30, 2024		4,846,635	​	$	10.40		8.24	​	$	527
Options exercisable at September 30, 2024		2,396,563	​	$	17.90		7.40	​	​	​
Options vested and expected to vest at September 30, 2024		4,846,635	​	$	10.40		8.24	​	$	527

Schedule of assumptions used to determine grant-date fair value of stock options

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​	​	​	​	​	​
​	​	Nine Months Ended September 30,			​
​		2024		2023	​
​	​	​	​	​	​
Expected volatility	​	92.99	%	90.37	%
Weighted average risk-free interest rate	​	3.98	%	4.05	%
Expected dividend yield	​	0.00	%	0.00	%
Expected term (in years)	​	6.46	​	6.48	​

Schedule of RSU activity under the 2016 Plan

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​	​	​	​	​	​
​	​	​	​	Weighted
​	​	​	​	Average Grant
​		Number of Shares		Date Fair Value
Outstanding at December 31, 2023		262,500	​	$	19.97
Vested	​	(27,500)	​	$	25.70
Forfeited	​	(7,500)	​	$	19.40
Outstanding at September 30, 2024		227,500	​	$	19.29

Schedule of recognized stock-based compensation expense related to the issuance of stock option awards and RSUs to employees and non-employees in the condensed consolidated statements of operations

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Three Months Ended September 30,

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Nine Months Ended September 30,

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2024

2023

2024

2023

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Research and development

$

720

​

$

737

​

$

2,076

​

$

2,086

General and administrative

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619

​

563

​

​

1,881

​

1,789

Total

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$

1,339

​

$

1,300

​

$

3,957

​

$

3,875

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Tabular disclosure of allocation of amount expensed and capitalized for award under share-based payment arrangement to statement of income or comprehensive income and statement of financial position. Includes, but is not limited to, corresponding line item in financial statement.

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Tabular disclosure of the number and weighted-average grant date fair value for restricted stock units that were outstanding at the beginning and end of the year, and the number of restricted stock units that were granted, vested, or forfeited during the year.

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Tabular disclosure for stock option plans. Includes, but is not limited to, outstanding awards at beginning and end of year, grants, exercises, forfeitures, and weighted-average grant date fair value.

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Tabular disclosure of the significant assumptions used during the year to estimate the fair value of stock options, including, but not limited to: (a) expected term of share options and similar instruments, (b) expected volatility of the entity's shares, (c) expected dividends, (d) risk-free rate(s), and (e) discount for post-vesting restrictions.

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