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CONDENSED CONSOLIDATED STATEMENTS OF CHANGES IN STOCKHOLDERS' EQUITY - USD ($)
$ in Thousands
Total
At-the-market Offering
Common stock
Common stock
At-the-market Offering
Additional paid-in capital
Additional paid-in capital
At-the-market Offering
Accumulated deficit
Beginning balance (shares) at Dec. 31, 2020     43,205,221        
Beginning balance at Dec. 31, 2020 $ 37,493   $ 4   $ 603,685   $ (566,196)
Increase (Decrease) in Stockholders' Equity [Roll Forward]              
Net loss (61,759)           (61,759)
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan (in shares)     305,105        
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan 393       393    
Stock-based compensation 6,748       6,748    
Issuance of common stock, net of issuance costs (in shares)     5,274,261 4,726,012      
Issuance of common stock, net of issuance costs 46,824 $ 29,187 $ 1   46,823 $ 29,187  
Ending balance (shares) at Sep. 30, 2021     53,510,599        
Ending balance at Sep. 30, 2021 58,886   $ 5   686,836   (627,955)
Beginning balance (shares) at Jun. 30, 2021     51,511,845        
Beginning balance at Jun. 30, 2021 74,893   $ 5   681,558   (606,670)
Increase (Decrease) in Stockholders' Equity [Roll Forward]              
Net loss (21,285)           (21,285)
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan (in shares)     50,754        
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan (10)       (10)    
Stock-based compensation 2,405       2,405    
Issuance of common stock, net of issuance costs (in shares)       1,948,000      
Issuance of common stock, net of issuance costs   2,883       2,883  
Ending balance (shares) at Sep. 30, 2021     53,510,599        
Ending balance at Sep. 30, 2021 $ 58,886   $ 5   686,836   (627,955)
Beginning balance (shares) at Dec. 31, 2021 53,577,744   53,577,744        
Beginning balance at Dec. 31, 2021 $ 48,636   $ 5   688,156   (639,525)
Increase (Decrease) in Stockholders' Equity [Roll Forward]              
Net loss (13,265)           (13,265)
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan (in shares)     202,635        
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan (3)       (3)    
Stock-based compensation $ 3,230       3,230    
Issuance of commitment shares (in shares) 1,667,593            
Issuance of common stock, net of issuance costs (in shares)       2,587,855      
Issuance of common stock, net of issuance costs   $ 1,471   $ 1   $ 1,470  
Ending balance (shares) at Sep. 30, 2022 58,035,827   58,035,827        
Ending balance at Sep. 30, 2022 $ 40,069   $ 6   692,853   (652,790)
Beginning balance (shares) at Jun. 30, 2022     58,005,616        
Beginning balance at Jun. 30, 2022 48,357   $ 6   691,682   (643,331)
Increase (Decrease) in Stockholders' Equity [Roll Forward]              
Net loss (9,459)           (9,459)
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan (in shares)     30,211        
Exercise of options, vesting of restricted stock units and shares issued under employee stock purchase plan (3)       (3)    
Stock-based compensation $ 1,174       1,174    
Ending balance (shares) at Sep. 30, 2022 58,035,827   58,035,827        
Ending balance at Sep. 30, 2022 $ 40,069   $ 6   $ 692,853   $ (652,790)