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Revenue (Tables) | 9 Months Ended
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Sep. 30, 2025
Revenue from Contract with Customer [Abstract] |
Schedule of Disaggregated Revenues | Our disaggregated revenues were as follows for the periods presented (in thousands):
|  | Three Months Ended September 30, |  |  | Nine Months Ended September 30, |
|  | 2025 |  |  | 2024 |  |  | 2025 |  |  | 2024 |
Revenues under collaborative agreements |  |  |  |  |  |  |  |  |  |  |  |
Upfront license fees (1) |  | $ | 1,762 |  |  | $ | 3,386 |  |  | $ | 5,564 |  |  | $ | 5,671 |
Other collaborative revenue (2) |  |  | 1,782 |  |  |  | — |  |  |  | 1,782 |  |  |  | — |
Total revenues under collaborative agreements |  |  | 3,544 |  |  |  | 3,386 |  |  |  | 7,346 |  |  |  | 5,671 |
Royalties, license and other revenues (3) |  |  | 137 |  |  |  | 393 |  |  |  | 602 |  |  |  | 960 |
Total revenue |  | $ | 3,681 |  |  | $ | 3,779 |  |  | $ | 7,948 |  |  | $ | 6,631 |
(1) All of the upfront license fee revenue recognized each period was included within deferred revenue as contract liabilities at the beginning of each period. (2) No ne of the other collaborative revenue recognized in 2025 was included within deferred revenues as contract liabilities as of the beginning of the three and nine months ended September 30, 2025. (3) Of the royalties, license and other revenues recognized each p eriod, $28,000 and $30,000 was included within deferred revenues as contract liabilities as of January 1, 2025 and 2024, respectively.
Schedule of Contract with Customer Contract Liability and Receivable | Accounts receivable, net, and deferred revenues (contra ct liabilities) from contracts with customers, including collaboration partners, consisted of the following (in thousands):
|  | September 30, 2025 |  |  | December 31, 2024 |
Accounts receivable, net - beginning of the year |  | $ | 638 |  |  | $ | 676 |
Accounts receivable, net - end of the period |  | $ | 127 |  |  | $ | 638 |
Contract liabilities |  |  |  |  |  |
Deferred revenues - beginning of the year |  | $ | 21,821 |  |  | $ | 29,501 |
Deferred revenues - end of the period |  | $ | 16,228 |  |  | $ | 21,821 |
Contract assets, which consist of revenue recognized and performance obligations satisfied or partially satisfied in advance of customer billing were $549,000 and zero as of September 30, 2025, and December 31, 2024, respectively, and was included within prepaid expenses and other current assets on the condensed consolidated balance sheet. There were no contract assets at the beginning of the nine months ending September 30, 2025 or the twelve months ending December 31, 2024.

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- Definition Tabular disclosure of receivable, contract asset, and contract liability from contract with customer. Includes, but is not limited to, change in contract asset and contract liability. + References Reference 1: http://www.xbrl.org/2009/role/commonPracticeRef -Topic 606 -SubTopic 10 -Name Accounting Standards Codification -Section 50 -Paragraph 10 -Publisher FASB -URI https://asc.fasb.org/1943274/2147479806/606-10-50-10 + Details
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- Definition Tabular disclosure of disaggregation of revenue into categories depicting how nature, amount, timing, and uncertainty of revenue and cash flows are affected by economic factor. + References Reference 1: http://www.xbrl.org/2009/role/commonPracticeRef -Topic 606 -SubTopic 10 -Name Accounting Standards Codification -Section 50 -Paragraph 5 -Publisher FASB -URI https://asc.fasb.org/1943274/2147479806/606-10-50-5 + Details
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