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Stock-Based Awards (Tables)
 9 Months Ended
Sep. 30, 2025
Share-Based Compensation Arrangement by Share-Based Payment Award [Line Items]  
Schedule of Stock Based Compensation Expense

Operating expenses within the condensed consolidated statements of operations include stock-based compensation expense as follows (in thousands):

 

 

 

Three Months Ended September 30,

 

 

Nine Months Ended September 30,

 

 

 

2025

 

 

2024

 

 

2025

 

 

2024

 

Research and development

 

$

179

 

 

$

172

 

 

$

531

 

 

$

476

 

General and administrative

 

 

1,020

 

 

 

1,158

 

 

 

3,123

 

 

 

3,286

 

Total stock-based compensation expense

 

$

1,199

 

 

$

1,330

 

 

$

3,654

 

 

$

3,762

 
Schedule of Anti-dilutive Securities Excluded from Computation of Earnings Per Share

The following common share equivalents were excluded from the computation of diluted net loss per common share for the periods presented because including them would have been antidilutive (in thousands):

 

 

 

Three and Nine Months Ended September 30,

 

 

 

2025

 

 

2024

 

Stock options

 

 

32,292

 

 

 

26,876

 

Restricted stock units

 

 

334

 

 

 

501

 

Warrants

 

 

41,097

 

 

 

—

 
2021 Equity Incentive Plan [Member]  
Share-Based Compensation Arrangement by Share-Based Payment Award [Line Items]  
Schedule of Share-based Compensation Activity

A summary of activity under the 2021 Plan is as follows (in thousands, except per share amounts):

 

 

 

Number
of Options
Outstanding
(in thousands)

 

 

Weighted
Average
Exercise Price
(per share)

 

 

Weighted
Average
Remaining
Contractual
Term (years)

 

 

Aggregate Intrinsic Value
(in thousands)

 

Balance at December 31, 2024

 

 

16,658

 

 

$

1.30

 

 

 

8.23

 

 

$

—

 

Options granted

 

 

7,823

 

 

$

0.56

 

 

 

 

 

 

 

Options exercised

 

 

(181

)

 

$

1.22

 

 

 

 

 

 

 

Options expired/forfeited/cancelled

 

 

(324

)

 

$

1.05

 

 

 

 

 

 

 

Balance at September 30, 2025

 

 

23,976

 

 

$

1.06

 

 

 

8.10

 

 

$

15,061

 

Options exercisable at September 30, 2025

 

 

10,148

 

 

$

1.34

 

 

 

7.16

 

 

$

3,570

 

Options exercisable and expected to vest
     at September 30, 2025

 

 

23,976

 

 

$

1.06

 

 

 

8.10

 

 

$

15,061

 

 

 

 

Number
of RSUs
Outstanding
(in thousands)

 

 

Weighted
Average Grant
Date Fair Value
(per share)

 

Balance at December 31, 2024

 

 

501

 

 

$

1.24

 

RSUs forfeited

 

 

(100

)

 

$

0.21

 

RSUs vested

 

 

(67

)

 

$

1.50

 

Balance at September 30, 2025

 

 

334

 

 

$

1.50

 

 
2012 Equity Incentive Plan and 2018 Inducement Option [Member]  
Share-Based Compensation Arrangement by Share-Based Payment Award [Line Items]  
Schedule of Share-based Compensation Activity

A summary of activity of the 2012 Plan, and the 2018 inducement option (which was issued to a Lineage executive outside of Lineage’s equity plans), is as follows (in thousands, except per share amounts):

 

 

 

Number
of Options
Outstanding
(in thousands)

 

 

Weighted
Average
Exercise Price
(per share)

 

Balance at December 31, 2024

 

 

10,068

 

 

$

1.84

 

Options exercised

 

 

(441

)

 

$

0.97

 

Options expired/forfeited/cancelled

 

 

(1,311

)

 

$

2.33

 

Balance at September 30, 2025

 

 

8,316

 

 

$

1.81