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Stock-Based Compensation (Tables) | 3 Months Ended
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Mar. 31, 2024
Summary of Assumptions Used in Black-Scholes Option Pricing Model | ​
​ | ​ | ​ | ​ | ​ | ​
​ | ​ | Three Months Ended | ​
​ | ​ | March 31, | ​
​ |  | 2024 |  | 2023 |
Risk-free interest rate |  | 4.12% | ​ | 3.62% - 4.22% |
Expected volatility |  | 89.6% | ​ | 76.3% - 81.3% |
Expected dividend yield |  | 0% | ​ | 0% |
Expected life of employee and Board options (in years) |  | 6.25 | ​ | 6.25 |
Summary of Stock Option Activity | ​
​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | ​ | ​ | Weighted | ​
​ | ​ | Number of | ​ | Average Exercise | ​
​ |  | Shares |  | Price |
Outstanding, December 31, 2023 |  | 7,897,647 | ​ | $ | 12.99 |
Granted |  | 1,875,125 | ​ |  | 0.99 |
Exercised |  | — | ​ |  | — |
Forfeited |  | (679,165) | ​ |  | 10.87 |
Expired |  | (277,075) | ​ |  | 13.36 |
Outstanding, March 31, 2024 |  | 8,816,532 | ​ | $ | 10.59 | ​
Options exercisable, March 31, 2024 |  | 4,827,562 | ​ | ​ | ​ | ​
Employees And Non-Employee Members Of Board Of Directors [Member] |
Summary of Restricted Stock Unit Activity | ​
​ | ​ | ​ | ​ | ​ | ​
​ | ​ | ​ | ​ | Weighted
​ | ​ | Number of | ​ | Average Grant
​ |  | Units |  | Date Fair Value
Outstanding, December 31, 2023 |  | 566,324 | ​ | $ | 9.27
Awarded |  | — | ​ |  | —
Vested and released |  | (186,375) | ​ |  | 11.04
Forfeited |  | (129,682) | ​ |  | 10.57
Outstanding, March 31, 2024 |  | 250,267 | ​ | $ | 7.28
Restricted stock units exercisable (vested and deferred), March 31, 2024 |  | — | ​ | ​ | ​
Employee Stock Option [Member] |
Summary of Compensation Expense Recognized | ​
​ | ​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | Three Months Ended | ​
​ | ​ | March 31, | ​
​ |  | 2024 |  | 2023 |
Research and development | ​ | $ | 504 | ​ | $ | 1,480 | ​
General and administrative | ​ |  | 1,229 | ​ |  | 1,241 | ​
Total stock option expense | ​ | $ | 1,733 | ​ | $ | 2,721 | ​
Restricted Stock Units [Member] |
Summary of Compensation Expense Recognized | ​
​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | Three Months Ended
​ | ​ | March 31,
​ |  | 2024 |  | 2023
Research and development |  | $ | 255 |  | $ | 179
General and administrative | ​ | ​ | 1,357 | ​ | ​ | 453
Total restricted stock unit expense |  | $ | 1,612 |  | $ | 632

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- Definition Tabular disclosure of allocation of amount expensed and capitalized for award under share-based payment arrangement to statement of income or comprehensive income and statement of financial position. Includes, but is not limited to, corresponding line item in financial statement. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Name Accounting Standards Codification -Topic 718 -SubTopic 10 -Section 50 -Paragraph 2 -Subparagraph (h)(1) -Publisher FASB -URI https://asc.fasb.org//1943274/2147480429/718-10-50-2 + Details
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- Definition Tabular disclosure of the number and weighted-average grant date fair value for restricted stock units that were outstanding at the beginning and end of the year, and the number of restricted stock units that were granted, vested, or forfeited during the year. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Name Accounting Standards Codification -Topic 718 -SubTopic 10 -Section 50 -Paragraph 2 -Subparagraph (c)(1) -Publisher FASB -URI https://asc.fasb.org//1943274/2147480429/718-10-50-2 + Details
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- Definition Tabular disclosure for stock option plans. Includes, but is not limited to, outstanding awards at beginning and end of year, grants, exercises, forfeitures, and weighted-average grant date fair value. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Name Accounting Standards Codification -Topic 718 -SubTopic 10 -Section 50 -Paragraph 2 -Subparagraph (c)(1) -Publisher FASB -URI https://asc.fasb.org//1943274/2147480429/718-10-50-2 Reference 2: http://fasb.org/us-gaap/role/ref/legacyRef -Name Accounting Standards Codification -Section 50 -Paragraph 2 -Subparagraph (d) -SubTopic 10 -Topic 718 -Publisher FASB -URI https://asc.fasb.org//1943274/2147480429/718-10-50-2 Reference 3: http://fasb.org/us-gaap/role/ref/legacyRef -Name Accounting Standards Codification -Section 50 -Paragraph 2 -Subparagraph (e) -SubTopic 10 -Topic 718 -Publisher FASB -URI https://asc.fasb.org//1943274/2147480429/718-10-50-2 + Details
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- Definition Tabular disclosure of the significant assumptions used during the year to estimate the fair value of stock options, including, but not limited to: (a) expected term of share options and similar instruments, (b) expected volatility of the entity's shares, (c) expected dividends, (d) risk-free rate(s), and (e) discount for post-vesting restrictions. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Topic 718 -SubTopic 10 -Subparagraph (f)(2) -Name Accounting Standards Codification -Paragraph 2 -Section 50 -Publisher FASB -URI https://asc.fasb.org//1943274/2147480429/718-10-50-2 + Details
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